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Lehigh University · Courses

BIOS

133 courses with the subject BIOS, each shown exactly as we captured it from the college's catalog, with every element we hold. Where the wording looks broken, that is our reading of the catalog, not the college's text.

BIOS 001
Subject
BIOS
Type
alias
Source
catalog.lehigh.edu
BIOS 010Bioscience in the 21st Century3

A multidisciplinary survey of advances in bioscience. Exploration of themebased topics (e.g., infectious diseases, cancer, genomebase medicine, engineered biomedical systems) coupled with social/ethical considerations. Three lectures per week. Participation in online multidisciplinary discussion, writing assignments, field trips, and/or other activities. Attribute/Distribution: NS, NW

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 032(EES 032) Oceanography3

An introduction to the structure, composition, and processes of the earth from a marine perspective. Topics include earth structure, plate tectonics, continental margins, coastal processes, seawater chemistry, ocean circulation, wave dynamics, primary productivity, plankton and plants, marine organisms and communities. May be combined with EES 022 or EES 004 for 4 credits. Attribute/Distribution: NS, NW

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 041Introduction to Cellular and Molecular Biology 03

Basic building blocks and higher order structures required for cellular processes. Topics include the character of membranes, the molecular/cellular basis of energy production, cell cycle progression, DNA replication, gene expression, basic Mendelian genetics, signal transduction, and cell division. Prerequisites: CHM 075 or CHM 025 or CHM 030 or CHM 040 Can be taken Concurrently: CHM 075, CHM 025, CHM 030, CHM 040 Attribute/Distribution: NS, NW

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 042Introduction to Cellular and Molecular Biology Laboratory1

Techniques and experiments related to the principal topics covered in BIOS 041; emphasis on experimental design and scientific communication. Prerequisites: BIOS 041 Can be taken Concurrently: BIOS 041 Attribute/Distribution: NS, NW, W

Subject
BIOS
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 043Phage Hunting Laboratory2

The first laboratory research course in a two-semester series (as part of the Howard Hughes Medical Institute's SEA-PHAGES Program) that focuses on the isolation (from local soil), as well as the physical and genomic characterization of novel bacteriophages that infect mycobacteria. Course will substitute for BIOS 42. Application and acceptance into Lehigh's SEA program required (see www.lehigh.edu/~insea); freshman status in the spring semester or permission of the instructor. Co-requisite: BIOS 41 or instructor permission. Prerequisites: BIOS 041 Can be taken Concurrently: BIOS 041 Attribute/Distribution: NS, Q

Subject
BIOS
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 044Introduction to Integrative and Comparative Biology 03

Experimental and historical approaches to the analysis of structural and functional properties in organisms. Use of scientific method to study species diversity. Introduction to the analysis of organismal attributes that explain behavioral repertoire and ecological relationships. Prerequisite: BIOS 041. Prerequisites: BIOS 041 Attribute/Distribution: CC, NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 045Introduction to Integrative and Comparative Biology Laboratory1

Experiments and discussions related to the topics covered in BIOS 044; emphasis on experimental design and scientific communication. Prerequisites: BIOS 044 Can be taken Concurrently: BIOS 044 Attribute/Distribution: LS, NS, Q

Subject
BIOS
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 091Special Topics1-4

Intensive study of a topic of special interest not covered in other courses. Consent of instructor required. Repeat Status: Course may be repeated. Attribute/Distribution: CC, NS, NW, W

Subject
BIOS
Credits (min)
1
Credits (max)
4
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 115Genetics 03

The structure, function, and continuity of hereditary information. Classical genetic analysis. Molecular biology of genes and genomes. Population genetics and evolution. Genetics of complex traits. Prerequisites: BIOS 041 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 116Genetics Laboratory1

Introduction to model organisms; techniques used in molecular genetics; experimental design and scientific communication. Prerequisites: BIOS 115 and (BIOS 042 or BIOS 043) Can be taken Concurrently: BIOS 115 Attribute/Distribution: LS, NS

Subject
BIOS
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 118Phage Genetics Laboratory2

Part of a 2-semester series focusing on genetic analysis of novel bacteriophage genomes to determine gene function using recombineering strategies. Phage genome annotation using bioinformatics for previously sequenced phage genomes. Additional genetics, molecular biology, and/or biochemical research on previously isolated bacteriophages may also be included. Prerequisites: BIOS 115 Can be taken Concurrently: BIOS 115 Corequisites: BIOS 115Attribute/Distribution: LS, NS, W

Subject
BIOS
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 130(MATH 130) Biostatistics 04

Elements of statistics and probability theory with emphasis on biological applications. Statistical analysis of experimental and observational data. Prerequisites: MATH 052 or MATH 022 or MATH 032 Attribute/Distribution: Q

Subject
BIOS
Credits (min)
4
Credits (max)
4
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 160Independent Study1-3

This independent study allows a student to explore a topic not covered in the existing curriculum under the direct supervision of an instructor. Repeat Status: Course may be repeated. Prerequisites: BIOS 041

Subject
BIOS
Credits (min)
1
Credits (max)
3
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 161Supervised Research1-3

Apprenticeship in ongoing faculty research program. Literature review, experimental design, data collection and analysis, and professional writing under faculty sponsor supervision. Only 3 credits can be counted toward any life science major. Consent of instructor required. Repeat Status: Course may be repeated. Prerequisites: BIOS 041 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
1
Credits (max)
3
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 191Special Topics1-4

Intensive study of a topic of special interest not covered in other courses. Consent of instructor required. Repeat Status: Course may be repeated. Attribute/Distribution: CC, NS, NW, W

Subject
BIOS
Credits (min)
1
Credits (max)
4
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 202Biomedical Externship0-3

Analysis of individualized experiences at external biomedical clinical or research sites. Limited enrollment. May not be taken for passfail grading. May not be used to satisfy any life science major or minor requirement. Consent of department chair required. Attribute/Distribution: NS

Subject
BIOS
Credits (min)
0
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 218SEA-GENES Laboratory2

A third semester extension of the SEA-PHAGES lab experience. A course-based research experience with goals established by the Howard Hughes Medical Institute - the cloning of all genes of a chosen bacteriophage, the testing of the functions of individual bacteriophage genes by phenotypic assays, the identification of host proteins that physically interact with a limited subset of bacteriophage proteins - the ultimate goal of which is establishing a complete data set for the chosen bacteriophage. Prerequisites: BIOS 043 and BIOS 118 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 231Entomology3

An exploration of the world of insects, the most diverse group of animals on Earth. Fundamentals of insect biology, including morphology, physiology, development, behavior, and ecology. Practical skills in insect collection, preservation, and identification. Topics will include the impact of insects on human society, including agriculture and public health. Prerequisites: BIOS 041 and BIOS 042 and BIOS 044 Attribute/Distribution: NW

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 234Comparative Vertebrate Anatomy4

A course in vertebrate zoology with emphasis on the study of homologous body structures in the various vertebrate classes and their relationship to the functional demands of habit and environment in each class. Detailed dissections of representative vertebrates are made in the laboratory. Two lectures and two laboratory periods. Prerequisites: BIOS 044 and BIOS 045 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
4
Credits (max)
4
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 235Human Physiology3

The goal of this course is to provide students with a thorough understanding of the major systems of the human body. Students will examine many of the major body systems including Nervous, Muscular, Cardiac, Respiratory, and Digestive systems. These systems will be analyzed by their components at both the cellular and molecular level. Students will also examine how each system functions as a whole and how it interacts with other systems at the organismal level. Prerequisites: BIOS 044 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 237(BIOC 237) Introductory Molecular Modeling and Simulation3

Key concepts, methods, and tools used in molecular modeling and simulation. A hybrid lecture/hands-on practice course using the lectures and tools in CHARMM-GUI (http://www.charmm-gui.org/lecture). Topics include (but not limited to) UNIX operating system, text editors, Python programming, scientific programming using Python, PDB (Protein Data Bank), molecular mechanics, minimization, molecular dynamics, Monte Carlo simulation. The understanding of these concepts and algorithms as well as their applications to well-defined practical examples involving currently important biological problems will be emphasized. Prerequisites: CHM 030 or CHM 040 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 238Epigenetics, Health, and Environment3

Foundational concepts in environmental epigenetics. Course will be centered around presentations and discussions of studies that address current issues, such as the mechanisms behind heritable health effects of acute malnourishment in human populations across multiple generations, or the potential for extremely fast evolutionary adaptation in species threatened by environmental change. This course may be listed as a Writing Intensive course. Prerequisites: BIOS 044 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 239Ecology and Evolution of Infectious Diseases3

In this course, we will explore the causes and consequences of infectious disease at multiple scales—from whole organisms to entire ecosystems. By evaluating primary literature, leveraging theoretical models, and considering experimental design, we will aim to apply evolutionary and ecological understanding to the epidemics of the past, present, and future. Prerequisites: BIOS 044 and BIOS 045 Attribute/Distribution: CC, NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 251Writing and Biological Sciences3

A course designed to acquaint students with some of the intellectual foundations of science, with attention to the distinctiveness of the biological sciences. Format includes readings, intensive writing, extemporaneous speaking, and discussion. May not be used to fulfill Biology B.A. elective requirements. Attribute/Distribution: NS, W

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 260Independent Study1-3

This independent study allows a student to explore a topic not covered in the existing curriculum under the direct supervision of an instructor. Repeat Status: Course may be repeated. Prerequisites: BIOS 041

Subject
BIOS
Credits (min)
1
Credits (max)
3
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 262Research Proposal3

Literature and methods of research in area of department faculty expertise. Requires development of detailed proposal for research to be performed in senior year. Must have major in any biological sciences degree program, junior standing, GPA of 3.0 in major, and consent of department. Attribute/Distribution: NS, W, WRIT

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 274(HMS 274) Neuroethics3

The intersection of neuroscience and ethics. History of biomedical science and current topics in neuroethics explored through weekly case studies and relevant readings in neurobiology. Examples include: definitions of mental illness, definitions of consciousness and brain death, addiction neuroscience, brain-machine interfaces, wearable technology, social determinants of health and equity within science and medicine. Reading and critical analysis of scientific articles, integration of biological concepts with moral reasoning, effective written communication and participation in peer review, oral presentations and group discussions. Prerequisites: BIOS 044 Attribute/Distribution: NS, W

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 276Central Nervous System and Behavior 03

Neuronanatomy and neurophysiology of animal and human behavior. Feeding, thirst, sleep, emotions, learning, and psychopathology. Prerequisites: BIOS 044 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 277Experimental Neuroscience Laboratory2

Structure and function of the mammalian brain with special attention to cellular morphology and organization. Widely used histological and behavioral techniques to determine how the shape and function of the nervous system regulates behavior. Experimental design, hypothesis testing, statistical analysis, reading and writing of scientific papers, basic histology and imaging. Prerequisites: BIOS 276 Can be taken Concurrently: BIOS 276 Attribute/Distribution: NS, W

Subject
BIOS
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 278Neurophysiology Laboratory2

The functional electrical underpinnings of the nervous system explored through direct recordings and thorough data analyses. Experimental design, hypothesis testing, numerical analysis, reading and writing of primary science. Prerequisites: BIOS 276 Can be taken Concurrently: BIOS 276 Attribute/Distribution: NS, Q

Subject
BIOS
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 279Experimental Molecular Neuroscience Laboratory2

Inquiry-based lab course emphasizing molecular and cellular neuroscience approaches to understanding the nervous system. Opportunity for making real research discoveries on the genetic and cellular underpinnings of brain function. Molecular genetic, bioinformatic, and neurochemical techniques. Can be taken more than once with instructor approval. Repeat Status: Course may be repeated. Prerequisites: BIOS 276 Can be taken Concurrently: BIOS 276 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 280Neurogenetics and Behavior Laboratory2

Inquiry-based lab course utilizing functional genetic and behavioral approaches to understand nervous system function. This course will provide students with opportunities to employ genome engineering techniques and behavioral analysis to make real research discoveries about the genetic underpinnings of behavior. This course is an approved lab for the Neuroscience major. Prerequisites: BIOS 115 and BIOS 116

Subject
BIOS
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 291Special Topics1-4

Intensive study of a topic of special interest not covered in other courses. Consent of instructor required. Repeat Status: Course may be repeated.

Subject
BIOS
Credits (min)
1
Credits (max)
4
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 300Apprentice Teaching1-4

Supervised participation in various aspects of the teaching of a course. Consent of instructor, department chairperson, and permission of the Dean required. Repeat Status: Course may be repeated.

Subject
BIOS
Credits (min)
1
Credits (max)
4
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 315Neuropharmacology3

Mechanisms of drug action in the central nervous system. Pharmacokinetics/pharmacodynamics. Depressant, stimulants, analgesics, and psychedelics. Treatments for neuropsychiatric disorders. Drug abuse. Prerequisites: BIOS 276 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 317Evolution 03

Mechanisms of evolution, emphasizing genetic structure and variation of populations, and isolation. Origin of species and higher taxa. Rates of evolution, extinction. Prerequisites: BIOS 115 Attribute/Distribution: CC, NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 318Advanced Phage Research Laboratory1-3

Apprentice-based laboratory research in phage biology in collaboration with faculty who direct Lehigh's SEA Phages Program (sponsored by the Howard Hughes Medical Institute). Research will focus on discoveries uncovered in SEA Phage courses and may include genetic, molecular, and biochemical analyses of novel bacteriophage genes, genomes, and novel gene functions. Additional research may include phage genome annotation using computational biology tools. Repeat Status: Course may be repeated. Prerequisites: BIOS 115 and BIOS 118 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
1
Credits (max)
3
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 320(ENTP 320) The Business of Life Science3

An examination of business process in startup, early stage and developing bioscience companies. Technology assessment, business plan and proposal preparation, financial strategies, resource management, intellectual property, and legal as well as regulatory issues. Cannot be used to fulfill major or minor requirements in Biological Sciences. Prerequisites: BIOS 121 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 323Evolution of Development3

This course examines how changes in the blueprint that describes the development of a multicellular animal from a single fertilized egg lead to the evolution of new species or new forms within a species. Prerequisites: BIOS 317 or BIOS 376 or BIOS 327 Can be taken Concurrently: BIOS 317, BIOS 376, BIOS 327

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 324Microbiology3

An examination of microbial life, including archaea, bacteria, fungi, protists and viruses. Emphasis on microbial molecular genetics and its relationship to the origin of life, human health/medicine, and the environment. Prerequisites: BIOS 115 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 325Microbiology Laboratory2

Laboratory studies of microorganisms, focusing on bacteria and fungi. Techniques for isolating, culturing, and identifying microorganisms. Experiments in microbial molecular genetics, phylogenetics, and evolution using traditional and modern techniques, as well as other topics covered in BIOS 324. Prerequisites: BIOS 324 Can be taken Concurrently: BIOS 324 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 326Coevolution3

Discussion-based seminar course covering readings from the primary literature. Course will examine evolutionary consequences of species interactions, interactions between the sexes, and genetic interactions. Topics will include predator-prey interactions, host-parasite interactions, sexual conflict, genomic conflict, mutualism, and more. Consequences of coevolution for biodiversity and human health will also be examined. Prerequisites: BIOS 317 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 327Development and Disease3

Development of organs from precursor cells; diseases that impact organ function. Focus on understanding how mutations or other causes influence organ development and function. Combination lecture and primary literature. Prerequisites: BIOS 376 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 328Immunology3

Distinction of "self" and "nonself" through humoral and cellular mechanisms. Antigens; biochemical structures, cellular mechanisms, genetic control and processing, phylogenetic distribution, diseased states. Prerequisites: BIOS 115 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 330Molecular Evolution3

This course will focus on evolution at the level of individual genes, proteins, and genomes, alternating between lecture and discussion of papers from the recent primary literature. Topics include pathways for adaptive evolution, directionality in evolution, epistasis, evolvability, genome rearrangements and speciation, gene duplication, and evolutionary dynamics. We will draw on examples of molecular evolution in nature, laboratory model systems, and human pathogens. Prerequisites: BIOS 317 or BIOS 345 Can be taken Concurrently: BIOS 317, BIOS 345 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 332Behavioral Neuroanatomy3

The study of neuroanatomy that underlies social and motivated behaviors and the techniques that support this study. Students learn by researching and reporting on original research, and ultimately designing and describing an original study on the topic. Prerequisites: BIOS 276 Attribute/Distribution: NS, W

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 334Species and Speciation3

Consideration of the origin of species. Discussion of a variety of "species" definitions and exploration of the evolutionary mechanisms by which new species arise. Alternation between lecture and discussion, drawing on the textbook and on current and classical literature. Prerequisites: BIOS 317

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 335(PSYC 335) Animal Behavior3

Discussion of the behavior of invertebrates and vertebrates and analysis of the physiological mechanisms responsible for behavioral stimuli, and adaptive value of specific behavior patterns. Prerequisites: BIOS 044 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 336Animal Behavior Laboratory2

Experiments and field observations illustrating principles discussed in BIOS 335. Emphasis on observing animals, performing experiments, collecting and analyzing data, and individual research. Six hours of laboratory per week. Prerequisites: BIOS 335 or BIOS 337 Can be taken Concurrently: BIOS 335, BIOS 337 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 337Behavioral Ecology3

Social systems of vertebrate and invertebrate groups. Emphasis on ecological and evolutionary factors that influence social behavior. Prerequisites: BIOS 044 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 338Neurodegenerative Diseases in Model Organisms3

Discussion-based seminar course on the use of model systems to investigate neurodegenerative diseases. The primary focus will be on invertebrate model systems, including Drosophila. Critical reading of primary scientific literature and student discussion is required. Prerequisites: BIOS 115 Attribute/Distribution: W

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 339Computational, Molecular Modeling and Simulation3

This course is designed to introduce the most basic and key concepts, methods, and tools used in molecular modeling and simulation. This class is a hybrid of lecture and hands-on practice styles, using the lectures and tools in CHARMM-GUI (http://www.charmm-gui.org/lecture). Some topics include the UNIX operating system, text editors, Python programming and Monte Carlo simulation. The understanding of these concepts and algorithms, as well as their applications to well-defined practical examples involving currently important biological problems will be emphasized. Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 340Molecular Basis of Disease3

Lectures and student projects on molecular mechanisms of human disease. Physiology of disease, molecular mechanisms, therapeutic approaches, ongoing research. Topics include: neurodegenerative diseases, cancer, autoimmune diseases, infectious diseases. Prerequisites: BIOS 115 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 341Living Together: The Science of Host-Microbe Interactions3

This course explores the fascinating and complex world of host-microbe interactions, examining the impact of host-associated microbes on biodiversity, ecology, and evolution. Using primary literature, we investigate the diversity of these interactions and the mechanisms driving them, and engage in lively group discussions and collaborative mini-reviews that hone scientific communication skills. By the end of the course, you will have gained a deep understanding of the latest research on host-microbe interactions and the tools to evaluate and communicate scientific ideas. Prerequisites: BIOS 044 or BIOS 121 and BIOS 115

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 342Cellular Basis of Human Disease3

Cell and molecular biological advanced topics relevant to human disease and/or health. Critical reading of the primary literature, discussion and student discussion required. Prerequisites: BIOS 367 or BIOS 411 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 343Personal Genomics3

Contemporary methods and technologies for investigating human genetic variation and its use for inferring ancestry and risk for disease, along with discussions of relevant policy and ethics. Readings will include primary scientific literature in population and statistical genetics, government publications, and news reports. Final projects will involve development of outreach and education resources in this topic for non-experts. Prerequisites: (BIOS 115 and BIOS 130) or BIOS 317 Attribute/Distribution: CC, NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 345Molecular Genetics3

The organization and replication of genetic material; mutagenesis; mechanisms of regulation; mechanisms of gene transmission involving prokaryotes and eukaryotes and their viruses; techniques for intervention into genetic organization and expression. Prerequisites: BIOS 115 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 346Molecular Genetics Laboratory2

Laboratory experiments related to the topics covered in BIOS 345. Emphasis is on molecular characterization of DNA and the principles of gene isolation and transfer. Prerequisites: BIOS 345 Can be taken Concurrently: BIOS 345 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 347Advanced Topics in Genetics3

Lectures and student projects on selected aspects of genetics such as the genetics and evolution of particular organisms, regulation of gene expression and transmission, human genetics, gene therapy, etc. Consent of department chair. Prerequisites: BIOS 115 and BIOS 116 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 348Marine Biology3

Ecology and adaptations of marine species, populations and ecosystems. Studying life in the ocean requires a perspective that spans spatial and temporal scales from the planetary to the microscopic, from the geologic to the physiological, and from the surface to the deepest seafloor. Through this course, students develop a fundamental understanding of how the physical environment of planet earth shapes and is shaped by marine organisms. Prerequisites: BIOS 041 and BIOS 042 Attribute/Distribution: CC, NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 349Molecular Ecology3

Molecular ecology is a field of research that seeks to answer questions in ecology, evolution, behavior and conservation; through the use of molecular tools. Biological scales range from organisms, to populations and species. This is a discussion-based course. Students read, present and analyze classic and modern scientific literature. Students also formulate a research proposal addressing outstanding questions in the field. Prerequisites: BIOS 044 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 353Virology3

An introduction to viruses and their interactions with host organisms. Topics include viral pathogenesis, from entry through release, viral immune evasion mechanisms, and viral evolution. Vaccines, antiviral strategies, as well as therapeutic uses of viruses for gene therapy will be emphasized. Prerequisites: BIOS 115 Attribute/Distribution: CC, NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 355The Brain Behind Behavior3

How brain mechanisms enable behavior is a central theme of neuroscience. This course gives a broad overview of the structure and function of major neural circuits (networks of interconnected nerve cells that work together to process and transmit information) that underlie various behaviors. We will start with gross neuroanatomy, followed by sensory and motor systems, innate behaviors (eat, drink, sleep, etc.) and cognitive behaviors (learning, memory, decision- making, language, etc.), and finally discuss consciousness and brain evolution. Prerequisites: BIOS 276

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 360Independent Study1-3

This independent study allows a student to explore a topic not covered in the existing curriculum under the direct supervision of an instructor. Repeat Status: Course may be repeated. Prerequisites: BIOS 041

Subject
BIOS
Credits (min)
1
Credits (max)
3
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 361Undergraduate Research1-3

Laboratory research under tutorial with a faculty member. Must have junior or senior standing. Consent of instructor required. Repeat Status: Course may be repeated.

Subject
BIOS
Credits (min)
1
Credits (max)
3
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 365Neurobiology of Sensory Systems3

The fundamental features of sensory systems in a diverse array of animals. Focus on how nervous systems detect, compute, and internally represent aspects of the environment from the single cell to whole system level. Special attention to the way sensory processing influences how we think about the biological basis of perception and possible mechanisms for consciousness. Instructor permission required. Prerequisites: BIOS 276 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 366Diseases of the Nervous System3

Neurobiological basis of CNS disorders, including affective, neurological and psychotic conditions. Emphasis on primary literature covering causes, diagnostic and treatment issues. Prerequisites: BIOS 276 or BIOS 382 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 367Cell Biology3

Molecular aspects of cell biology. Emphasis on membrane structure and function, organelle biogenesis, cell motility, the cytoskeleton, and extracellular matrix. Prerequisites: BIOS 115 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 368Cell Biology Laboratory2

Basic methods used in cell biology laboratories around the world and the opportunity to carry out an independent research project. Techniques include histology and microscopy (both white and fluorescent light), tissue culture and sterile procedures, cellular fractionation, nuclear import assays, and immunological probing. Consent of department required. Prerequisites: BIOS 367 Can be taken Concurrently: BIOS 367 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 371(CHM 371) Elements of Biochemistry I 03

A general study of carbohydrates, proteins, lipids, nucleic acids and other biological substances and their importance in life processes. Protein and enzyme chemistry are emphasized. Must have completed one year of organic chemistry. Prerequisites: (CHM 031 or CHM 041) and CHM 110 and CHM 112 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 372(CHM 372) Elements of Biochemistry II3

Dynamic aspects of biochemistry; enzyme reactions including energetics, kinetics and mechanisms; metabolism of carbohydrates, lipids, proteins and nucleic acids; photosynthesis, electron transport mechanisms, coupled reactions, phosphorylations, and the synthesis of biological macromolecules. Prerequisites: BIOS 473 or ((BIOS 371 or CHM 371) and BIOS 041) Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 375Methods in Developmental Biology Lab2

Detection of gene expression and protein expression in vivo or in vitro. Mutants and/or transgenics examined. Students address research questions of instructor. Prerequisites: BIOS 376 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 376Developmental Biology3

Differentiation of multicellular organisms from a single cell. Axis determination; gradients; induction and pattern formation viewed through modern analysis of regulated gene expression. Lecture topics on organ formation paired with discussions on birth defects and human diseases. Prerequisites: BIOS 115 and (BIOS 116 or BIOS 118) Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 377(CHM 377) Biochemistry Laboratory 03

Laboratory studies of the properties of chemicals of biological origin and the influence of chemical and physical factors on these properties. Laboratory techniques used for the isolation and identification of biochemicals. Prerequisites: (BIOS 371 or CHM 371) and (BIOS 041) Can be taken Concurrently: BIOS 371, CHM 371 Attribute/Distribution: ND

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 381Physical Biochemistry3

Topics include: thermodynamics of biological systems; Forces acting on and between biological molecules; Principles of macromolecular structure; Physical methods used to characterize biomolecules; and other topics to be determined. Prerequisites: (BIOS 371 or CHM 371) and (BIOS 041) Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 382(PSYC 382) Endocrinology3

Organization and function of endocrine and neuroendocrine systems in regulating physiology and behavior. Emphasis on mammalian systems. Focus on critical thinking and problem-solving with reference to basic literature. Prerequisites: BIOS 044 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 383Biological Sciences Colloquia 01

Analysis of weekly colloquia in the biological sciences. Repeat Status: Course may be repeated. Prerequisites: BIOS 044 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
1
Credits (max)
1
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 384Eukaryotic Signal Transduction3

Signal transduction between cells of multicellular eukaryotic organisms examined in the context of specialized functions that include: nutrition, hormones and neurotransmitters, vision, muscle contraction, adhesion, and the immune system. The evolution of cancer based on mutations in these signaling systems. Prerequisites: BIOS 367 or BIOS 372 or CHM 372 or BIOS 382 or BIOS 365 Attribute/Distribution: W

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 385Synapses, Plasticity and Learning3

Communication between neurons. Physiology of synaptic transmission; varying forms of neuronal plasticity; acquisition, encoding, and retrieval of memory. Prerequisites: BIOS 276 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 386Genes and the Brain3

Modern molecular genetics techniques applied to complex brain processes. Emphasis on DNA and RNA manipulation strategies to elucidate mechanisms of complex behaviors. Animal models of learning, behavioral plasticity, and neuropsychiatric diseases. Prerequisites: BIOS 276 Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 387Biological Sciences Honors Seminar1

Development, presentation and implementation of research proposals, and discussions of research. Required for senior biology, molecular biology, biochemistry, and behavioral neuroscience majors pursuing departmental honors. Departmental permission required. Attribute/Distribution: ND

Subject
BIOS
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 388Biological Sciences Honors Seminar1

Continuation and extension of BIOS 387. Departmental permission required. Attribute/Distribution: ND

Subject
BIOS
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 389Honors Project1-6

Repeat Status: Course may be repeated.

Subject
BIOS
Credits (min)
1
Credits (max)
6
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 391Special Topics1-4

Intensive study of a topic of special interest not covered in other courses. Must have junior or senior standing. Consent of instructor required. Repeat Status: Course may be repeated. Attribute/Distribution: ND

Subject
BIOS
Credits (min)
1
Credits (max)
4
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 393Thesis3

Literature review and design of project in selected area, execution of the project, final report and presentation. Consent of department required. Intended for senior majors in BIOS only. Consent of instructor required. Repeat Status: Course may be repeated. Attribute/Distribution: W

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 401Professional Skills for Biological Sciences Graduate Students3

Students learn expectations and fundamental skills related to success in the biological sciences. The course is designed to help students make the most out of their graduate education. Students learn the principles underlying fundable, publishable research, and how these general principles can be applied to their specific research area. They learn to write and review manuscripts and grant proposals by serving on a mock editorial board and scientific review panel. They gain experience in giving oral presentations. Readings are from texts on scientific writing and research styles, and from original journal articles and grant proposals written by the faculty. Required of all Integrative Biology graduate students.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 402Ethics and Rigor in Research1

This course covers material critical to the responsible conduct of science, in addition to data reproducibility, rigor, research ethics, and misconduct.

Subject
BIOS
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 403Methods and Logic in Biological Research3

Methods and Logic is a first-semester paper-based course for graduate students that presents classic papers in biology with a particular emphasis on the on the logical reasoning and design of experiments that led to major breakthroughs.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 404(PSYC 404) Behavioral Neuroscience3

Theoretical and empirical issues in biopsychology. Must have graduate standing.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 405Special Topics in Molecular Biology1-3

Research, conferences, and reports on selected topics not covered in the general graduate offerings. Repeat Status: Course may be repeated.

Subject
BIOS
Credits (min)
1
Credits (max)
3
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 406Biological Sciences Seminar 01

An advanced seminar in current developments including departmental research. Required for candidates for graduate degrees in molecular biology. Repeat Status: Course may be repeated.

Subject
BIOS
Credits (min)
1
Credits (max)
1
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 407Research in Biological Science1-9

Laboratory investigations in one of the department's research areas. Repeat Status: Course may be repeated.

Subject
BIOS
Credits (min)
1
Credits (max)
9
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 408Responsible Conduct of Science0

Responsible practice in research. Training in general laboratory methods; human subjects concerns; radiation safety; chemical hazards; aseptic technique; physical, mechanical, biological, and fire hazards; animal welfare. Occupational and workplace considerations. Recombinant DNA guidelines; patent and proprietary rights; controversies over applications of science. Appropriate aspects required of investigators in all departmental research projects.

Subject
BIOS
Credits (min)
0
Credits (max)
0
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 410Special Topics in Behavioral and Evolutionary Bioscience1-3

Readings and discussions on selected topics not covered in the general graduate offerings.

Subject
BIOS
Credits (min)
1
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 411Advanced Cell Biology3

Cell structure and biochemistry, as related to specialized cell functions.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 415Synapses, Plasticity and Learning3

Communication between neurons. Physiology of synaptic transmission; varying forms of neuronal plasticity; acquisition, encoding and retrieval of memory.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 421Molecular Cell Biology I3

Molecular aspects of cell structure, cell motility, intracellular transport; and biomembrane dynamics. Prerequisites: BIOS 411

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 422Molecular Cell Biology II3

Molecular aspects of gene expression, including genome structure and replication, RNA synthesis/processing, and protein synthesis. Prerequisites: BIOS 345 or BIOS 345

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 423Evolution of Development3

This course examines how changes in the blueprint that describes the development of a multicellular animal from a single fertilized egg lead to the evolution of new species or new forms within a species. Prerequisites: BIOS 317 or BIOS 376 or BIOS 327 Can be taken Concurrently: BIOS 317, BIOS 376, BIOS 327

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 424Advanced Neurobiology of Sensory Systems3

This course is designed to provide an overview of core principles of neuroscience through exploration of sensory systems. The course will provide an intensive review of fundamental neural signaling followed by a broad introduction to the major sensory pathways. Focus will be on major organizing principles of neural systems, and information processing. Student discussions and presentations will incorporate current literature and concepts.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 426Coevolution3

Discussion-based seminar course covering readings from the primary literature. Course will examine evolutionary consequences of species interactions, interactions between the sexes, and genetic interactions. Topics will include predator-prey interactions, host-parasite interactions, sexual conflict, genomic conflict, mutualism, and more. Consequences of coevolution for biodiversity and human health will also be examined.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 427Techniques in Cell and Molecular Biology3

Laboratory experiences in three or more cell and molecular biological techniques: gel electrophoresis of nucleic acids/proteins; polymerase chain reaction; DNA/RNA sequencing; molecular hybridization techniques; fluorescence microscopy; confocal microscopy; flow cytometry; electron microscopy tissue preparation; immunological detection methods; molecular cloning techniques; oocyte microinjection techniques; tissue culture methods; and autoradiography.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 428Molecular Evolution3

Evolution at the level of individual genes, proteins, and genomes. Lectures and discussion of papers from the recent primary literature. Topics include pathways for adaptive evolution, directionality in evolution, epistasis, evolvability, genome rearrangements and speciation, gene duplication, and evolutionary dynamics. Examples of molecular evolution in nature, laboratory model systems, and human pathogens. Prerequisites: BIOS 317 or BIOS 345 Can be taken Concurrently: BIOS 317, BIOS 345

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 431Advanced Topics in Cell Biology3

Current research problems in cell biology. Repeat Status: Course may be repeated. Prerequisites: BIOS 367 or BIOS 367 or BIOS 411

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 432Advanced Topics in Molecular Genetics3

Current research in molecular genetics. Repeat Status: Course may be repeated.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 433Advanced Topics in Developmental Biology3

Current research problems in developmental biology. Repeat Status: Course may be repeated. Prerequisites: BIOS 345 or BIOS 345

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 434Speciation3

Discussion-based seminar course covering readings from classical and current literature, including both theoretical and empirical contributions. Topics will be included species concepts, reproductive isloation, mechanisms and modes of speciation, and current approaches to studying speciation.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 435Epigenetics3

Foundational concepts in environmental epigenetics. Course will be centered around presentations and discussions of studies that address current issues, such as the mechanisms behind heritable health effects of acute malnourishment in human populations across multiple generations, or the potential for extremely fast evolutionary adaptation in species threatened by environmental change.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 438Neurodegenerative Diseases in Model Organisms3

Discussion-based seminar course on the use of model systems to investigate neurodegenerative diseases. The primary focus will be on invertebrate model systems, including Drosophila. Critical reading of primary scientific literature and student discussion is required. Prerequisites: BIOS 115

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 439Advanced Behavioral Ecology3

Critical evaluation of the theoretical foundation in sociobiology. Emphasis placed on kinship, altruism, mate choice, parental investment, parent-offspring conflict, etc. Lectures and seminars. Not open to students who have taken BIOS 337. Prerequisites: BIOS 317 or BIOS 317

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 441Living Together: The Science of Host-Microbe Interactions3

This course explores the fascinating and complex world of host-microbe interactions, examining the impact of host-associated microbes on biodiversity, ecology, and evolution. Using primary literature, we investigate the diversity of these interactions and the mechanisms driving them, and engage in lively group discussions and collaborative mini-reviews that hone scientific communication skills. By the end of the course, you will have gained a deep understanding of the latest research on host-microbe interactions and the tools to evaluate and communicate scientific ideas.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 442Cellular Basis of Human Disease3

Cell and molecular biological advanced topics relevant to human disease and/or health. Critical reading of the primary literature, discussion and student discussion required. Prerequisites: BIOS 367 or BIOS 411

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 443Personal Genomics3

Contemporary methods and technologies for investigating human genetic variation and its use for inferring ancestry and risk for disease, along with discussions of relevant policy and ethics. Readings will include primary scientific literature in population and statistical genetics, government publications, and news reports. Final projects will involve development of outreach and education resources in this topic for non-experts.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 448Marine Biology3

Ecology and adaptations of marine species, populations, and ecosystems. With an average depth of more than two miles, the ocean makes up more than 95% of the habitable space on our planet. Ocean ecosystems are essential life support systems for the entire earth. Studying life in the ocean requires a perspective that spans spatial and temporal scales from the planetary to the microscopic, from the geologic to the physiological, and from the surface to the deepest seafloor. Through this course,.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 449Molecular Ecology3

Molecular ecology is a field of research that seeks to answer questions in ecology, evolution, behavior and conservation; through the use of molecular tools. Biological scales range from organisms, to populations and species. This is a discussion-based course. Students read, present and analyze classic and modern scientific literature. Students also formulate a research proposal addressing outstanding questions in the field.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 450Developmental Neurobiology3

Fundamental mechanisms underlying neural development. Early events leading to the induction of the neuroectoderm and the reorganization of the vertebrate central nervous system during adulthood and aging. Major developmental events such as phenotype commitment, cell migration, differentiation and growth cone guidance. Emphasis on the interplay between concepts emerging from organismal and molecular levels of analyses.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 453General Neuroanatomy3

Graduate level study of the neuroanatomy and neurochemistry of systems that underlie behavior in vertebrates. Emphasis will be on the traditional and novel methodologies used to reveal neuroanatomical pathways as well as the function of these pathways. Consent of department required.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 455The Brain Behind Behavior3

This course gives a broad overview of the structure and function of major neural circuits (networks of interconnected nerve cells that work together to process and transmit information) that underlie various behaviors. We will start with gross neuroanatomy, followed by sensory and motor systems, innate behaviors (eat, drink, sleep, etc.) and cognitive behaviors (learning, memory, decision- making, language, etc.), and discuss consciousness and brain evolution.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 457Advanced Behavioral Neuroendocrinology3

A seminar course that covers current primary literature on the hormone-nervous system interactions that underlie physiology and behavior. The course covers the neuroendocrinology of reproduction, sex behavior, parental behavior, social behavior, agonistic and territorial behavior, learning and memory, homeostasis (caloric, nutritional, water and salt balance, temperature regulation), circadian rhythms and seasonality in a variety of vertebrates.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 460Independent Study1-3

This independent study allows a student to explore a topic not covered in the existing curriculum under the direct supervision of an instructor. Repeat Status: Course may be repeated.

Subject
BIOS
Credits (min)
1
Credits (max)
3
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 464Molecular Biology of Eukaryotic Organisms3

Comparative analysis of several eukaryotes as model systems in cell biology, developmental biology, genetics, and molecular biology.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 466Structure and Function of RNAs and Ribonucleoprotein Complexes3

Biochemistry and function of small nuclear RNPs, RNase P, ribosomes, self-splicing introns, signal recognition particle, RNA viruses. Functions of RNA in DNA replication, in regulation, as an enzyme, and as a repressor.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 471Eukaryotic Signal Transduction3

Signal transduction between and within cells of multicellular organisms examined in the context of specialized functions that include: nutrition, hormones and neurotransmitters, vision, muscle contraction, adhesion and the immune system. The evolution of cancer based on mutations in these signaling systems. Lecture, discussion, and student presentations. Prerequisites: (BIOS 372 or CHM 372 or BIOS 411)

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 472(CHM 472) Lipids and Membranes3

The study of lipids and lipid membranes similar to those found in mammalian cells including methods of synthesis, surface activity, bilayer and micellar structures, lipid mixing, fluidity, permeability and membrane stability. Special emphasis will be given to the current evidence for and against the lipid raft hypothesis. Prerequisites: BIOS 372 or CHM 372

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 473(CHM 473) Principles of Biochemistry I3

Study of proteins, carbohydrates, lipids, nucleic acids and other biological substances. Protein and enzyme chemistry are emphasized. Must have completed one year each of general chemistry and organic chemistry.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 474Computational, Molecular Modeling and Simulation3

This course is designed to introduce the basic and advanced concepts, methods, and tools used in molecular modeling and simulation. This class is a hybrid of lecture and hands-on practice styles, using the lectures and tools in CHARMM-GUI (http://www.charmm-gui.org/lecture). Topics include (but are not limited to) the UNIX operating system, text editors, Python programming, scientific programming using Python, PDB (Protein Data Bank) format, molecular mechanics, minimization, molecular dynamics, Monte Carlo simulation. The understanding of these concepts and algorithms, as well.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 477(CHM 477) Topics in Biochemistry1-3

Selected areas of biochemistry, such as mechanisms of enzyme action, new developments in the chemistry of lipids, nucleic acids, carbohydrates and protiens. Repeat Status: Course may be repeated.

Subject
BIOS
Credits (min)
1
Credits (max)
3
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 483Special Topics in Behavioral Neuroscience3

Examination of the biological substrates of behavior. Topics may include animal communication, sociobiology, behavioral endocrinology, or behavior genetics. Repeat Status: Course may be repeated.

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 486Genes and the Brain3

Modern molecular genetics techniques applied to complex brain processes. Emphasis on DNA and RNA manipulation strategies to elucidate mechanisms of complex behaviors. Animal models of learning, behavioral plasticity, and neuropsychiatric diseases. Attribute/Distribution: NS

Subject
BIOS
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 488Seminar in Neuroscience, Behavior, and Evolution1

Advanced seminar in current research developments.

Subject
BIOS
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 490Thesis1-6

Repeat Status: Course may be repeated.

Subject
BIOS
Credits (min)
1
Credits (max)
6
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu
BIOS 499Dissertation1-15

Repeat Status: Course may be repeated.

Subject
BIOS
Credits (min)
1
Credits (max)
15
Credit unit
Credits
Type
course
Repeatable
may be repeated.
Edition
2026-2027
Catalog
2026-27 Catalog
Source
catalog.lehigh.edu

Source: Lehigh University's catalog, linked per course · table learning_unit · CourseShelf publish 59