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Cornell University · Courses

BIOEE

65 courses with the subject BIOEE, each shown exactly as we captured it from the college's catalog, with every element we hold. Where the wording looks broken, that is our reading of the catalog, not the college's text.

BIOEE 1130The Art and Science of Birds2

Art and science have always been fundamentally linked. Before the invention of photography, illustration was the main documentation tool in science; even now, artistic representations are used to communicate important scientific discoveries. In this course students learn the basics of bird anatomy in tandem with sketching techniques to foster an appreciation of how science and art can reinforce each other in enriching one's life and study. Led by the staff biological illustrator at the Cornell Lab of Ornithology, students begin with the fundamentals of observational drawing to improve accuracy before moving onto watercolors. Interwoven into art lessons are scientific lectures produced by Cornell ornithologists, blending art with current science to cover a range of interconnected topics.

Subject
BIOEE
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 1150Techniques of Avian Specimen Preparation2

This course has two main objectives: 1) to illustrate the diverse uses of natural history collections for research, teaching and conservation, and 2) to introduce students to the fine art of avian specimen preparation. Students will learn multiple specimen preparation techniques (skeletons, spread wings, and round skins) and will be required to prepare 20 round skins to receive course credit. Students will be evaluated on the quality of their specimens and on their participation in an in-class debate surrounding the value of natural history collections.

Subject
BIOEE
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 1180Evolution3

Evolution is the central unifying concept in biology. This course examines evolution as a science and places it in an historical context. Classes focus on descent with modification, natural selection, evolutionary genetics, the history of the earth, the information content of the fossil record, and diversification processes. The science of evolutionary biology is presented in the context of a broader history of ideas in science. The course also explores the importance of evolutionary thinking in the 21st century, including antibiotic and pesticide resistance, personalized genomics, human evolution, and evolutionary ecology. Courses of Study: Intended for students with no background in college biology.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 1540Introductory Oceanography3

This class relies more on intuitive reasoning rather than complicated mathematical formulas to convey basic concepts about how the ocean works. For this reason, the class is very accessible to non-science majors. The class covers standard material about how the ocean works, but also includes current environmental threats facing the ocean such as global warming, ocean acidification, overfishing and coastal pollution. Students will gain a depth of knowledge about the ocean and global warming to enable them to speak and write confidently about contemporary public issues regarding the health of the ocean, global warming and a sustainable future. This course satisfies the Physical and Biological Sciences (PBS) requirement for students in most colleges. For students in A&S and CALS, this course counts as in-college credit.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 1560Introductory Oceanography with Laboratory4

This class relies more on intuitive reasoning rather than complicated mathematical formulas to convey basic concepts about how the ocean works. For this reason, the class is very accessible to non-science majors. The class covers standard material about how the ocean works, but also includes current environmental threats facing the ocean such as global warming, ocean acidification, overfishing and coastal pollution. Students will gain a depth of knowledge about the ocean and global warming to enable them to speak and write confidently about contemporary public issues regarding the health of the ocean, global warming and a sustainable future. This course satisfies the Physical and Biological Sciences (PBS) requirement for students in most colleges and the Introductory Life Sciences/Biological Sciences requirement for students in CALS. For students in A&S and CALS, this course counts as in-college credit. This course is suitable for non-life sciences majors.

Subject
BIOEE
Credits (min)
4
Credits (max)
4
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 1610Introductory Biology: Ecology and the Environment3-4

This course provides an introduction to ecology, covering interactions between organisms and the environment at scales of populations, communities, and ecosystems. Ecological principles are used to explore the theory and applications of major issues facing humanity in the 21st century, including population dynamics, disease ecology, biodiversity and invasive species, global change, and other topics of environmental sustainability.

Subject
BIOEE
Credits (min)
3
Credits (max)
4
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 1640FWS: Topics in Ecology and Evolutionary Biology3

The First-Year Writing Seminar provides the opportunity to write extensively about issues related to ecology and evolutionary biology. Topics vary by section.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 1760Biostatistics with R programming language3

This course in Biostatistics uses the R programming language as a platform for analysis. Students will be introduced to different types of statistical analysis while becoming comfortable writing basic code in the R programming language. Topics will include: descriptive statistics, inference, experimental design and hypothesis testing; assumptions behind statistical models and choice of statistical tests; analysis of variance and covariance; general linear models and interactions; regression; and parametric and non-parametric tests, bootstrapping, and permutation testing. No prior statistical or programming experience is required.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 1780An Introduction to Evolutionary Biology and Diversity4-5

Considers explanations for pattern of diversity and the apparent good fit of organisms to the environment. Topics include the diversity of life, the genetics and developmental basis of evolutionary change, processes at the population level, evolution by natural selection, modes of speciation, long-term trends in evolution, origin of humans.

Subject
BIOEE
Credits (min)
4
Credits (max)
5
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 1781Introduction to Evolution and Diversity4

Considers explanations for pattern of diversity and the apparent good fit of organisms to the environment. Topics include the diversity of life, the genetics and developmental basis of evolutionary change, processes at the population level, evolution by natural selection, modes of speciation, long-term trends in evolution, origin of humans.

Subject
BIOEE
Credits (min)
4
Credits (max)
4
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 2150Advanced Techniques of Avian Specimen Preparation2

This course builds upon skills introduced in BIOEE 1150 (Techniques of Avian Specimen Preparation) and offers students opportunities to improve and diversify specimen preparation skills and become more involved in CUMV activities. Students will gain experience preparing a diversity of birds (e.g., waterbirds, long-necked birds, big birds, long-legged birds), many of which require species-specific preparation techniques. Students will also improve skills in identifying key anatomical structures for aging birds, evaluating reproductive status, documenting appropriate metadata for modern scientific specimens, and further hone skills in discussing the values of natural history collections.

Subject
BIOEE
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 2233Field Lectures in Biodiversity and Evolution1

A surprisingly diverse, beautiful, and intriguing set of species live in our backyard. This class will meet outdoors for 3 hours per week, rain or shine, with extensive walking in natural areas (forests, fields, streams, agricultural lands). Field lectures will be on the specific plants and animals we encounter, discussing their ecology and exploring the evolutionary relationships between species. The class is hands-on, students can expect to get dirty and must be comfortable in the field. Connections will be drawn between theory learned in the classroom and real world biology and biodiversity out of doors. We will experience habitats and species, take notes, and write about biodiversity.

Subject
BIOEE
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 2525Ecology and Conservation of Wildlife in the Neotropics2

This on-campus and international field course combination provides participating students with a broad introduction to the research process in field ecology, with literature and hands-on examples drawn from the fauna and flora of coastal Patagonia (Argentina), an area that provides us with unprecedented access to both marine and terrestrial wildlife as well as exposure to conservation challenges and success stories. The course begins in the latter part of the Fall semester (BIOEE 2525) when it meets twice weekly for seven weeks, largely to discuss relevant papers from the scientific literature with an emphasis on best practices in designing field studies to address questions in Neotropical ecology, behavioral ecology, conservation, and evolutionary biology. During the 2+ week field component in January (BIOEE 2526), students travel among field sites in Patagonia and put this knowledge to work in an experiential context by designing and implementing a series of research projects, including numerous short 'blitz' projects and several longer, more intensive independent projects; many of these field studies involve close-hand observations of marine mammals, penguins, or other seabirds. The course (BIOEE 2527) is focused on building skills in data analysis and scientific writing, based on the data collected in the field by each student.

Subject
BIOEE
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 2526Ecology and Conservation of Wildlife in the Neotropics II1

This on-campus and international field course combination provides participating students with a broad introduction to the research process in field ecology, with literature and hands-on examples drawn from the fauna and flora of coastal Patagonia (Argentina), an area that provides us with unprecedented access to both marine and terrestrial wildlife as well as exposure to conservation challenges and success stories. The course begins in the latter part of the Fall semester (BIOEE 2525) when it meets twice weekly for seven weeks, largely to discuss relevant papers from the scientific literature with an emphasis on best practices in designing field studies to address questions in Neotropical ecology, behavioral ecology, conservation, and evolutionary biology. During the 2+ week field component in January (BIOEE 2526), students travel among field sites in Patagonia and put this knowledge to work in an experiential context by designing and implementing a series of research projects, including numerous short 'blitz' projects and several longer, more intensive independent projects; many of these field studies involve close-hand observations of marine mammals, penguins, or other seabirds. The course (BIOEE 2527) is focused on building skills in data analysis and scientific writing, based on the data collected in the field by each student.

Subject
BIOEE
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 2527Neotropical Wildlife Biology1

This on-campus and international field course combination provides participating students with a broad introduction to the research process in field ecology, with literature and hands-on examples drawn from the fauna and flora of coastal Patagonia (Argentina), an area that provides us with unprecedented access to both marine and terrestrial wildlife as well as exposure to conservation challenges and success stories. The course begins in the latter part of the Fall semester (BIOEE 2525) when it meets twice weekly for seven weeks, largely to discuss relevant papers from the scientific literature with an emphasis on best practices in designing field studies to address questions in Neotropical ecology, behavioral ecology, conservation, and evolutionary biology. During the 2+ week field component in January (BIOEE 2526), students travel among field sites in Patagonia and put this knowledge to work in an experiential context by designing and implementing a series of research projects, including numerous short 'blitz' projects and several longer, more intensive independent projects; many of these field studies involve close-hand observations of marine mammals, penguins, or other seabirds. This course (BIOEE 2527) is focused on building skills in data analysis and scientific writing, based on the data collected in the field by each student.

Subject
BIOEE
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 2641Tropical Field Ornithology II1.5
Subject
BIOEE
Credits (min)
1.5
Credits (max)
1.5
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 2642Analysis and Interpretation of Bird Observations1
Subject
BIOEE
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 2740The Vertebrates: Form, Function, and Evolution4

This course in organismal biology explores the anatomy and function of vertebrates from an evolutionary perspective. Lectures cover the origin, anatomy, physiology, ecology, and evolution of vertebrate groups, with a focus on organ systems, life history strategies, locomotion, behavior, ecology, and conservation. A concurrent lab provides hands-on experience with vertebrate anatomy and morphology. Overall, students will examine how form relates to function across diverse vertebrate lineages, investigate patterns of convergent evolution, and analyze factors driving biodiversity differences among groups. This course prepares students for advanced vertebrate biology courses (Ichthyology, Herpetology, Ornithology, Mammalogy) and provides excellent preparation for pre-vet and pre-med students through its comparative anatomical and physiological approach to understanding the organization of the vertebrate body.

Subject
BIOEE
Credits (min)
4
Credits (max)
4
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 3233Advanced Field Lectures in Biodiversity3

A surprisingly diverse, beautiful, and intriguing set of species live in our backyard. This class will meet outdoors for 4 hours per week, rain or shine, with extensive walking in natural areas (forests, fields, streams, agricultural lands). Field lectures will be on the specific plants and animals we encounter, discussing their ecology and exploring the evolutionary relationships between species. The class is hands-on, students can expect to get dirty and must be comfortable in the field. Connections will be drawn between theory learned in the classroom and real world biology and biodiversity out of doors. We will experience habitats and species, take notes, and write about biodiversity. Discussions and assignments will support oral and written communication about how to study biodiversity.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 3250Evolutionary Medicine3

This course focuses on how evolutionary principles can help us better understand health and disease. Concepts from evolutionary biology (e.g., life history theory, coevolution, conflict, constraints and trade-offs) will be applied to key problems in medicine and public health, with case studies on antibiotic resistance, aging, cancer, autoimmune disease, and why pathogenic organisms cause harm to their hosts.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 3550Ecological and Environmental Data Analytics3

Ecology and Environmental Science have entered an era of increasing data volume and complexity, demanding rigorous quantitative reasoning and effective data communication. This course advances data literacy of undergraduate and graduate students by covering theoretical foundations and practical coding skills to analyze, interpret, and present ecological and environmental data. Through integrated lectures and laboratory sessions, students will gain knowledge and hands-on experience for data curation, hypothesis testing, regression modeling, and visualization. The course also introduces new machine learning and AI tools for ecological data analytics. Students will develop proficiency in Python and R using real-world data sets spanning organismal to landscape scales. Designed for students in biology and environmental science, this course builds a rigorous, modern quantitative toolkit for ecological research and environmental problem-solving.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 3610Advanced Ecology4

This course provides an in-depth survey of ecology emphasizing conceptual foundations and the integration of experimental and quantitative approaches, including physiological ecology, population and community ecology, ecosystem biology, and ecological modeling. Current and classical ecological research is used to introduce major concepts and methods, derive major ecological principles, and critically discuss their applicability on multiple organizational levels, on multiple scales, and in a variety of ecosystems. Lab sections focus on discussion, experimental techniques, and computational skills.

Subject
BIOEE
Credits (min)
4
Credits (max)
4
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 3611Field Ecology3

Exercises designed to give students direct experience with field research to address ecological hypotheses, with emphasis on developing observational skills and basic methods in population and community ecology. Topics include methods in plant succession, niche relationships, influence of herbivores and competitors on plant communities, aquatic food web analysis, use of scientific collections, and presenting research results in written and oral form. We will visit a diversity of habitats and natural areas in Central New York. Students will conduct an independent research project and present their findings.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 3690Chemical Ecology3

Why are chilies so spicy? This course examines the chemical basis of interactions between species and is intended for students with a basic knowledge of chemistry and biology. Focuses on the ecology and chemistry of plants, animals, and microbes. Stresses chemical signals used in diverse ecosystems, using Darwinian natural selection as a framework. Topics include: plant defenses, microbial warfare, communication in marine organisms, and human pheromones.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 3730Biodiversity and Biology of the Marine Invertebrates3

Introduction to the biology, natural history, and evolution of the major invertebrate phyla, concentrating on marine representatives. In addition to the evolution of form and function, lectures cover aspects of ecology, behavior, physiology, chemical ecology, and current research. The discussion section will focus on current research papers with marine invertebrates.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 3731Methods in Marine Invertebrate Biodiversity1

BIOEE3731 provides an immersive experience working in a research laboratory focused on understanding the evolution of biodiversity in marine invertebrates. Participants will perform experiments to investigate the relationships between DNA sequence diversity, cell and tissue morphology, and animal evolution. Students will learn several specialized research techniques including fluorescent microscopy, scanning electron microscopy, genome editing (CRISPR/Cas9), microinjection, and immunohistochemistry. Experiments will be conducted in the Babonis Lab (E142 Corson Hall), and learning will be assessed by completion of a semester-long research project. Because of the intensive hands-on nature of the course, only a small group of students will be accepted. Acceptance is by interview only.

Subject
BIOEE
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 3780Digital Morphology through CT1

This course is an introduction to CT visualization for its applications in comparative biology of the vertebrates. Students will learn and practice the exploration of vertebrate anatomy with OSIRIX 3-D visualization software or its future replacement; work on student-designed projects and/or a large survey of the vertebrates based on CT scans from specimens in the Cornell museum as well as the Smithsonian and other museums around the world.

Subject
BIOEE
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4000Ecology and Evolution of Infectious Diseases3

This course introduces students to the field of infectious disease ecology, an area of study that has developed rapidly over the past three decades and addresses some of the most significant challenges to human health and conservation. Students will learn about the incredible diversity of parasitic organisms, arguably the most abundant life forms on the planet, and examine how pathogens invade and spread through host populations. Throughout the course, an emphasis will be placed on understanding of infectious diseases dynamics at the population level, and on quantitative approaches for studying pathogen spread and impacts. Specific topics include types of pathogens and their ecological properties, epidemiology and impacts on host populations, types of transmission, evolution of resistance and virulence, drivers of the emergence of new diseases, parasites in the context of ecological communities, strategies for controlling outbreaks, and the role of parasites in biodiversity and conservation.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4460Plant Behavior and Biotic Interactions, Lecture3

How do plants respond to antagonists, such as herbivores and pathogens? What are the checks and balances that keep mutualist organisms in their tight interactions? How are symbioses organized on molecular, metabolic and ecological levels? What are the molecular, plant hormonal, and metabolic mechanisms mediating plant biotic interactions with other organisms? What ecological and evolutionary consequences do these interactions have for the fitness of the plants and their interactors? This course provides an overview of plants' myriad interactions with antagonists and mutualists, from microbes to multicellular organisms, and explains the underlying ecological and evolutionary concepts. It gives an introduction to the study of induced plant responses in the light of a behavioral biology framework.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4461Plant Behavior and Biotic Interactions, Laboratory1

Laboratory course covering topics presented in BIOEE 4460/BIONB 4460/PLSCI 4460.

Subject
BIOEE
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4500Mammalogy, Lectures and Laboratory4

This course examines the diversity and biology of mammals from an evolutionary perspective through an integrated lecture, laboratory and field experience. Students gain a foundational understanding of mammalian biology by exploring origins, evolution, diversity, biogeography, physiology, ecology, life history, and behavior, with emphasis on the defining traits of mammals and their variation across modern lineages. Laboratories complement lectures through hands-on study of specimens in the Cornell University Museum of Vertebrates. Field trips in New York provide opportunities to observe wild mammals, practice field identification, and apply methods in population biology. By integrating conceptual frameworks with specimen-based laboratories and field observations, students develop a deeper understanding of mammalian diversity and the evolutionary and ecological processes that shape it.

Subject
BIOEE
Credits (min)
4
Credits (max)
4
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4530Speciation: Genetics, Ecology, and Behavior4

Advanced course in evolutionary biology focusing on the pattern and process of speciation and the nature of origin of behavioral, morphological, physiological and ecological traits that form the intrinsic barriers to gene exchange. Lecture topics include species concepts and definitions, the history of ideas about speciation, the biological basis of intrinsic barriers to gene exchange, current models for the origin of such barriers, genetic architecture of speciation, rates of speciation. Emphasis is on developing a rigorous conceptual framework for discussing speciation and on detailed analysis of a series of case histories.

Subject
BIOEE
Credits (min)
4
Credits (max)
4
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4550Insect Ecology4

You will learn to think like an ecologist by studying the fundamental principles of insect ecology and the types of questions ecologists ask, seeing how ecology can be used to understand and solve environmental problems, and putting this knowledge into action during group activities in the lab and field.

Subject
BIOEE
Credits (min)
4
Credits (max)
4
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4560Stream Ecology4

Lecture examines patterns and processes in stream ecosystems, including geomorphology and hydrology, watershed-stream interactions, trophic dynamics, biogeochemistry, disturbance, and conservation and management. Field and laboratory exercises focus on experimental and analytical techniques used to study stream ecosystems, including techniques to measure stream discharge, physical habitat, water chemistry, and stream biota. Field project with lab papers.

Subject
BIOEE
Credits (min)
4
Credits (max)
4
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4570Limnology: Ecology of Lakes, Lectures3

Limnology is the study of inland freshwaters, which includes lakes, ponds, reservoirs, rivers, streams, and wetlands. This course focuses on lakes and ponds, and examines their physical, chemical, and biological properties. A lake's physical (e.g., size, light, temperature, mixing) and chemical (elements and compounds) structure directly affects aquatic organisms in terms of species abundance, diversity, and interactions. Organisms in turn influence their physical and chemical environment. This class will introduce students to fundamental concepts in limnology, which we will build upon to examine real-world challenges facing freshwater ecosystems (e.g., eutrophication, invasive species, climate change).

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4571Limnology: Ecology of Lakes, Laboratory2

Laboratories and field trips devoted to studies of the biological, chemical, and physical properties of lakes and other freshwater environments. Exercises focus on understanding the freshwater environment, on experimentation, and on understanding ecological processes within lakes.

Subject
BIOEE
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4610Ecological and Evolutionary Statistics3

This course develops quantitative and programming skills essential for designing, analyzing, and communicating ecological and evolutionary research. Emphasis is placed on sampling design, hypothesis formulation, statistical inference, and data analysis as an integrated process rather than separate steps. Students will learn to translate biological questions into testable predictions, evaluate them with data, and interpret results critically. Instruction focuses on conceptual understanding and informed choice of methods, preparing students for independent learning of advanced techniques. Data analysis is conducted in R. Case studies draw from contemporary ecological and evolutionary research. The course is tailored for students pursuing an Honors thesis, and the in-class project allows students to analyze data directly related to their Honors research. Grades are based on written assignments, final projects, and in-class participation.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4660Physiological Ecology, Lectures3

Detailed survey of how physiological processes help organisms meet the challenges of their environment. Lectures explore physiological adaptation, thermal biology, respiration and oxygen transport, resource acquisition and allocation, water use and limitation, energetics and nutrition, hormones, stress physiology, and developmental physiology. Readings draw from the primary literature and textbooks.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4661Physiological Ecology, Laboratory2

Detailed survey of the physiological approaches used in understanding the relationships between organisms and their environment. Laboratories apply physiological techniques to specific ecological problems and cover aspects of experimental design and computer-aided data analysis.

Subject
BIOEE
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4690Food, Agriculture, and Society3

Multidisciplinary course dealing with the social and environmental impact of food production in the United States and developing countries. Agroecosystems of various kinds are analyzed from biological, economic, and social perspectives. The impacts of traditional, conventional, and alternative agricultural technologies are critically examined in the context of developed and developing economies. Specific topics include biodiversity and ecosystem services in agriculture, transgenic crops, land use for energy production, urban agriculture, and sustainable development.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4700Herpetology, Lectures3

Lectures cover various aspects of the biology of amphibians and reptiles, including evolution, zoogeography, ecology, behavior, and physiology.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4750Ornithology, Lectures3

This course covers many aspects of avian biology, including ecology, behavior, evolution, anatomy, physiology, and conservation. This is an active learning-style offering in which students complete pre-class work through the Cornell Lab of Ornithology's Bird Academy online learning platform, and then engage in interactive activities and exercises during class periods. BIOEE 4750 may be taken alone, or students can choose to enroll simultaneously in the associated laboratory classes, BIOEE 4751.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4751Ornithology, Laboratory1

This laboratory course covers many aspects of avian biology, including ecology, behavior, evolution, anatomy, physiology, and conservation. Some laboratory sessions will involve dissection of avian material, and the study of skeleton and scientific study skin specimens. Other sessions will involve outdoor activities including observations of avian behavior and demonstrations of standard ornithological field techniques, including methods of capturing, handling, and banding birds.

Subject
BIOEE
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4752Ornithology, Worldwide Avian Diversity Laboratory2

Birds present a wondrous diversity of behaviors, ecological lifestyles, and appearances. This course provides an in-depth survey of the natural history and evolutionary relationships of all Orders and Families of living birds from across the globe. Cornell is the home of the Birds of the World online resource which provides background information on worldwide avian diversity, and BIOEE 4752 also makes extensive use of the avian specimen materials of the Cornell University Museum of Vertebrates, where the weekly lab sessions are held.

Subject
BIOEE
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4760Ichthyology: Biology of Fishes, Lectures3

Introduction to the systematic study of fossil and living fishes: their anatomy, physiology and functional morphology, behavior, ecology, diversity, evolution, classification, and identification. Emphasizes marine fishes from different habitats (temperate, tropical coral reef, intertidal, and deep sea), local freshwater species, and interesting freshwater fishes from around the world, especially South America, Africa and Australia.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4761Ichthyology: Biology of Fishes, Laboratory1

Laboratory course covering topics presented in BIOEE 4760. Two field trips, including one full-day weekend trip required. Live animals are studied in the field and are sometimes used in the laboratory for nondestructive demonstrations or experiments. The systematics and dissection laboratories use preserved specimens.

Subject
BIOEE
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4762The Biology, Evolution, and Lives of Fishes3

Fishes and fishing are linked throughout the history of humans. While early fishing was assuredly for survival, its modern form includes both subsistence and sport. This course will dive into the target of fishing- the most diverse vertebrate group; the fishes. Together we will explore numerous aspects about the biology of fishes including the diversity, evolution, anatomy, ecology, behavior, physiology, and conservation of the fishes of the world. Cornell is a world leading institution in the study of vertebrate biology, and has a long history of training students in the importance of specimen-based research. This is a college level course that will utilize the methods from many scientific disciplines in our approaches to studying fishes, though the only prerequisite is a strong interest in fishes.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4780Ecosystem Biology and Global Change4

Examines ecosystem structure, carbon and energy flow, and nutrient cycles, and how these processes respond to environmental variations. Reviews classic ecosystem experiments, and considers effects on ecosystems from human-driven environmental changes, such as climate change, air pollution, excess nitrogen use, and land-use change.

Subject
BIOEE
Credits (min)
4
Credits (max)
4
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4790Paleobiology4

Surveys the major groups of invertebrate organisms and their evolutionary histories, and the theoretical and practical principles of paleontology, from biostratigraphy to macroevolution. Intended to fill out the biological backgrounds of Earth and atmospheric science students concerning the nature and significance of the fossil record for their respective studies, and the paleontological backgrounds of biology students interested in ecology and evolution.

Subject
BIOEE
Credits (min)
4
Credits (max)
4
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4800Ecological Genetics4

Ecological Genetics focuses on the application of population genetic concepts in ecological contexts, with emphases on measuring adaptation in natural populations, detecting the effects of population demography, and determining the genetic basis of quantitative traits. Illustrative examples will be drawn from the primary research literature to demonstrate experimental techniques and methods of data analysis on single-gene, multi-locus and genome-wide scales.

Subject
BIOEE
Credits (min)
4
Credits (max)
4
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4940Special Topics in Ecology and Evolutionary Biology1-6

The department offers trial courses or seminars under this number. Offerings vary by semester. The same course is not to be offered more than twice under this number. For 2022-2023 descriptions, please go to the department website.

Subject
BIOEE
Credits (min)
1
Credits (max)
6
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 4980Teaching Experience1-4

Designed to give qualified undergraduate students teaching experience through actual involvement in planning and assisting in biology courses. This experience may include supervised participation in a discussion group, assisting in a biology laboratory, assisting in field biology, or tutoring.

Subject
BIOEE
Credits (min)
1
Credits (max)
4
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 6000Ecology and Evolution of Infectious Diseases3

This course introduces students to the field of infectious disease ecology, an area of study that has developed rapidly over the past three decades and addresses some of the most significant challenges to human health and conservation. Students will learn about the incredible diversity of parasitic organisms, arguably the most abundant life forms on the planet, and examine how pathogens invade and spread through host populations. Throughout the course, an emphasis will be placed on understanding of infectious diseases dynamics at the population level, and on quantitative approaches for studying pathogen spread and impacts. Specific topics include types of pathogens and their ecological properties, epidemiology and impacts on host populations, types of transmission, evolution of resistance and virulence, drivers of the emergence of new diseases, parasites in the context of ecological communities, strategies for controlling outbreaks, and the role of parasites in biodiversity and conservation.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 6550Ecological and Environmental Data Analytics3

Ecology and Environmental Science have entered an era of increasing data volume and complexity, demanding rigorous quantitative reasoning and effective data communication. This course advances data literacy of undergraduate and graduate students by covering theoretical foundations and practical coding skills to analyze, interpret, and present ecological and environmental data. Through integrated lectures and laboratory sessions, students will gain knowledge and hands-on experience for data curation, hypothesis testing, regression modeling, and visualization. The course also introduces new machine learning and AI tools for ecological data analytics. Students will develop proficiency in Python and R using real-world data sets spanning organismal to landscape scales. Designed for students in biology and environmental science, this course builds a rigorous, modern quantitative toolkit for ecological research and environmental problem-solving.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 6602Graduate Field Course in Ecology3

Designed to give graduate students experience in defining questions and designing field investigations. The course is based at the Archbold Biological Station in central Florida over spring break and during the following week. The class visits several ecosystems including sand pine scrub, cattle ranches, cypress swamps, and the Everglades.

Subject
BIOEE
Credits (min)
3
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 6900Seminar in Ecology and Evolution of Infectious Diseases1

Graduate-level discussion of the ecology, epidemiology, genetics, and evolution of infectious disease in animal and plant systems. Weekly discussion of research papers published in the primary scientific literature. Participation in discussion and presentation of at least one paper required for course credit.

Subject
BIOEE
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 7500Limnology Seminar2

This course is a discussion of current topics in limnology. Students will have the opportunity to present and receive feedback on their research in limnology. Topics include (but are not limited to) biogeochemistry, community ecology, ecosystem ecology, microbial ecology, and population ecology in inland freshwaters.

Subject
BIOEE
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 7600Special Topics in Evolution and Ecology1-3

Lectures and intensive study of special topics in the field of Ecology & Evolutionary Biology. Content varies each semester.

Subject
BIOEE
Credits (min)
1
Credits (max)
3
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 7640Plant-Insect Interactions Seminar1

Group intensive study of current research in plant-insect interactions. Topics vary from semester to semester but include chemical defense, coevolution, insect community structure, population regulation, biocontrol, tritrophic interactions, and mutualism.

Subject
BIOEE
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 7650Professional Development in Ecology and Evolutionary Biology2

Group discussion on professional activities for academic ecologists and evolutionary biologists. Topics include: writing a CV, publication strategies, finding a postdoc, the tenure process, grant writing, manuscript reviews, networking, public speaking, and scientific collaboration. Active “present, critique and revise” sessions will allow participants to improve written materials and oral presentations through peer and instructor feedback.

Subject
BIOEE
Credits (min)
2
Credits (max)
2
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 7670Current Topics in Ecology and Evolutionary Biology4.5

Critical evaluation and discussion of theory and research in ecology and evolutionary biology. Lectures by faculty and student-led discussions of topics in areas of current importance.

Subject
BIOEE
Credits (min)
4.5
Credits (max)
4.5
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 7720Generative AI for Research: Tools and Judgment1

This seminar examines how generative AI can be integrated into graduate research in ways that are effective, transparent, and intellectually responsible. Designed as a discussion-centered forum, the course explores practical uses of generative AI for coding, literature synthesis, data analysis, and writing, alongside broader questions about judgment, authorship, skill development, academic integrity, and the role of human expertise. Readings, case studies, and tool demonstrations will support weekly discussion. Readings, case studies, and short tool demonstrations will support weekly discussion. The overall goal is to help students develop a thoughtful personal framework for when and how generative AI should, and should not, be used in their own research practice, while cultivating an adaptable approach that can evolve as AI capabilities, norms, and expectations continue to change.

Subject
BIOEE
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 7800Graduate Seminar in Ornithology1

Group intensive study of current research in ornithology built around a research seminar series covering a wide variety of projects and topics in ornithology and related disciplines, including avian ecology, evolution, conservation, behavior, and physiology. Each weekly seminar is followed by informal discussion and networking that builds connections among the large community of Cornell students and scholars with an interest in avian biology. All students with an interest in this subject area are welcome and deep knowledge of ornithology is not expected or required. Instructor permission is not required for enrollment.

Subject
BIOEE
Credits (min)
1
Credits (max)
1
Credit unit
Credit
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 8990M.S. Thesis Research1-15

Thesis research conducted by an M.S. student in the field of ecology and evolutionary biology with advice and consultation of a major professor who is a member of the field.

Subject
BIOEE
Credits (min)
1
Credits (max)
15
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu
BIOEE 9990Ph.D. Dissertation Research1-15

Dissertation research conducted by a Ph.D. student in the field of ecology and evolutionary biology with advice and consultation of a major professor who is a member of the field.

Subject
BIOEE
Credits (min)
1
Credits (max)
15
Credit unit
Credits
Type
course
Edition
2026-2027
Catalog
2026-2027 Catalog
Source
catalog.cornell.edu

Source: Cornell University's catalog, linked per course · table learning_unit · CourseShelf publish 61