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Keystone College · Courses

BIOL

124 courses with the subject BIOL, each shown exactly as we captured it from the college's catalog, with every element we hold. Where the wording looks broken, that is our reading of the catalog, not the college's text.

BIOL 1008General Biology I: Focuses on the

scientific method, cell biology, genetics, evolution, and biotechnology. Provides the background to become a knowledgeable, literate, and active participant in an increasingly technological society. Focuses on ethical considerations of modern biological issues. For the non-science major; not open to students who have passed BIOL 1125 or

Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1009Human Health Issues: A one3

semester general biology course designed for the non-major. Focuses on human anatomy, physiology, diseases and health care maintenance. Provides the necessary background to become an active consumer of contemporary health care. This class is for the non-science major. Spring, online only, 3 hours lecture and discussion,

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1010Introduction to Field2

Biology: Provides a field-oriented experience with concentrated study of natural history, ecology, and conservation. Emphasizes introducing students to a number of field, laboratory, and classroom skills to prepare them for upper-level field courses and internships. Students should be prepared to experience the outdoor environment under all weather conditions. Fall, 2 hours lecture, discussion, fiel work.

Subject
BIOL
Credits (min)
2
Credits (max)
2
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1013Fall Wildflowers: Investigates the1

common wild flowering plants (herbs, shrubs, and trees) of the northeastern United States. Familiarizes students and prepares students to identify many common species and families and appreciate the status of Pennsylvania’s endangered plants. Students prepare museum quality herbarium specimens of collected plants Fieldwork required,

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1014Amazing Mosses: Examines the1

biology, morphology and ecology of mosses found in the northeastern United States. Features the investigation of bryophytes in the field and lab. Students should expect to spend time in the lab developing their microscopy skills. Students gain an understanding of how to identify the common species found in Northeastern Pennsylvania. A weekend day trip is required. Fieldwork required,

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1015Insects: An introductory course that1

covers the biology and ecology of insects. Discusses where to find insects, how to collect and preserve them and, through the use of various keys, how to identify them. A collection of insects will be done by each student. Fieldwork required,

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1019Geology of Northeast1

Pennsylvania: Journey through deep time, exploring ancient environments by examining the deposits left behind. Covers field techniques to identify rocks, fossils, and a multitude of 198 Keystone College geologic structures. Observations from the field will be used to reconstruct the ancient depositional environment. Classes consist of mixed lectures and field trips to local geologic outcrops. A Saturday field trip is required where students will transect Pennsylvania from south to north, studying the rock formations along the way. Some possible field trip stops include “the Whale Back” and Centralia. Fieldwork required,

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1023Forest Stewardship: Introduces1

principles and methods of forest stewardship including the philosophy of forest stewardship, identification, and ecology of some common trees, forest resource measurements, timber management, and wildlife management. Fieldwork required,

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1028Invasive Species: Addresses the1

d ecological and physical impacts of invasive exotic species in northeastern Pennsylvania, focusing on identification, culture and invasion scenarios that degrade native plant communities in urban, agricultural and wetland environments. The identification portion of the course utilizes the Woodlands Campus as a study area. Includes an afternoon/early evening field trip to a nearby degraded wetland, woodland, and farm. Fieldwork required,

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1038Marine Mammals: Focuses on whales1

dolphins, porpoises, and seals. Following land- based introductory sessions that dive into anatomy, taxonomy, and natural history, students travel to the Atlantic coast to see these magnificent animals in their natural habitat. Fieldwork required,

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1043Winter Ecology: Examines how plants1

and animals survive the rigors of cold and dryness during Pennsylvania’s winters. Studies the important impacts of snow cover on the lives of wildlife. Dress warmly, because this class spends many of its meeting dates from January through March in the outdoors. Fieldwork required,

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1048Forest Health: Identification of non1

native insects who continue to pose a significant threat to Pennsylvania forest ecosystems. Examines native and non-native insects and their ecological impact to forest composition. Focuses on insect biology, life cycles, host plants, and management strategies. Fieldwork required,

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1049Ecology of Vernal Pools1

Examines the biology and chemistry of vernal pools. Vernal pools fill with rainwater in the spring but often dry out in summer. During that time, they are alive with salamanders, frogs, tadpole, shrimp and many other creatures. Weekend field trip required; travel fee determined at time of travel. Fieldwork required,

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1052Lichens: Introduces the world1

of lichens, identification, biology, and ecology. Examines parts of the lichen used to identify these complex organisms. Discusses the general types of lichens as well as their habitats. Collect common lichens found in this area. Discusses lichens’ role in the ecology of their habitats, as well as human uses, and utilizes keys to identify the various species. Fieldwork required,

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1055Biology and Geology of Caves

Studies the formation and conservation of caves, as well as the organisms that call them home. Features a weekend-long caving trip to the limestone caverns of upstate New York. Travel fee determined at time of travel, 1 credit.

Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1058The Living Susquehanna

Experience the story of the Susquehanna River through a two-day kayak trip from Meshoppen to Falls. Learn about the longest river on the east coast. Covers the geology of the watershed, history, industrial uses, health of the river, and looking to the future. Travel fee determined at time of travel, 1 credit.

Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1061Mushrooms and Other Fungi1

Observe the various types of mushrooms. These characteristics will be used to identify the species of mushrooms found. Note: this is not a wild edibles course. Discusses poisonous and non- poisonous species. Fieldwork required,

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1073Spiders and Arachnids

Introductory course covers the biology and ecology of spiders and their kin (mites, ticks, harvesters). Field course that discusses where to find spiders, how to collect and preserve them, and how to identify them. Spider webs, egg cocoons, their food, courtship and anatomy will be discussed. Students will collect arachnids and requires one night session for nocturnal collections. Fieldwork required, 1 credit. 199 Keystone College

Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1077Winter Botany: Identify the many1

flowering plants by the skeletons that remain standing during the winter. Teaches important anatomical features of the flowers and fruits of the common wildflowers of the area. Time in the field and classroom will utilize keys to identify the various species, an important skill for wetlands workers. Fieldwork required,

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1078Understanding Lyme Disease1

Examines what individuals need to know about preventing, understanding, and treating Lyme disease and tick-borne illnesses. Focuses on how to be tick smart, and the latest advancements available to take back our life if affected by Lyme disease.

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1079Fossils of NEPA: Hunt for fossils that1

once thrived in ancient swamps over 350-million years ago. Introduces the Carboniferous Period, also known as the coal- bearing era, and the unique plants and animals that existed then. Collect fossils and examine how these extinct plants formed into anthracite coal and the marine organisms which pressurized into limestone. Search for plants fossils, anthracite coal, minerals, trilobites, crinoids, brachiopods and other ocean animals. Learn how to identify and categorize while contributing findings to Keystone’s fossil and mineral collection. Students will be able to keep many of the fossil finds as well. Visit two coal mine tours, which offer railroad rides deep into actual anthracite coal mines. Note: An all-day Saturday and Sunday weekend field collecting trip is required. Fieldwork required,

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1081Wildlife Forensics: Introduces1

principles and methods of criminal and civil investigations of wildlife crimes. Topics include international wildlife trade, laboratory methods, and the involvement of law enforcement. Connections made between conservation and forensic efforts. Classes include guest speakers, case studies, and mock investigations.

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1082Preserving Honeybees: Hands- on1

course that investigates honeybee colony management techniques and environmental practices to promote bee health and protect bees from disease. Create a nucleus colony from larger hives and design a natural setting in which honeybees can thrive. Work in groups to conduct research on bee-related topics of interest. 1 hour lecture, discussion, hands-on experiences.

Subject
BIOL
Credits (min)
1
Credits (max)
1
Credit unit
credit
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1120Environmental Science: Studies3

human interactions with the environment. Focuses on ecological principles of ecosystem structure, population dynamics of humans and other organisms. Considers sources and supplies of biological, food, soil, mineral and energy resources, as well as methods of mitigating the effects of their use and exploitation. The philosophy of sustainable development will underlie the course, as will the role of the individual in the decision-making process. Corequisite: BIOL 1121. Spring, 2 hours lecture and discussion, 2-hour laboratory,

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1125General Biology I: Introduces4

science majors to topics such as the nature of science, basic chemistry, cell structure and function, reproduction, and basic genetics. Dissection required. Corequisite: BIOL 1126. Fall, Spring, and Summer, 3-hour lecture and discussion, 3-hour laboratory,

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1130General Biology II: Introduces4

science majors to topics such as population biology, ecology, evolution, energy relationships, and the diversity of non-plant kingdoms. Dissection required. Corequisite: BIOL 1131. Fall, Spring and Summer, 3-hour lecture and discussion, 3-hour laboratory,

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 1125 with a “C” or better.
BIOL 1135Introduction to Forensic Science

I: Includes scientific and investigative techniques of various forensic disciplines including scene investigation, documentation, photography, patterns evidence, firearms, fingerprints, forensic arson, and forensic entomology. Examines the role of the expert witness and the scientific method in forensic investigation at the crime scene. Corequisite:

Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1136Fall, lecture, 3-hour3

laboratory, 4 credits.

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
hours
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1140Medical Terminology: Studies the3

technical and specialized anatomical, surgical, pharmaceutical, and common medical phrases. Fall and Spring (online only), 3 hours lecture and discussion,

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1145Online only (not offered every3

semester), 3 credits lecture and discussion,

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 1150Botany: Introduces science majors to4

the structure, function, and diversity of plants as well as algae. Studies morphology, anatomy, physiology, taxonomy, ecology, and economic botany.

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 1130 with a “C” or better. Corequisite: BIOL 1151. Dissection required. Fall, 3 hours lecture and discussion, 3 hour laboratory,
BIOL 1501Special Topics - Problem Solving for3

the Sciences: Properly acclimates incoming first- year students to mathematical problem solving that is relevant to the sciences in the aim of better facilitating their success in first year course work such as general chemistry and general biology. 1 hour lecture, discussion, hands-on assignments, 1 credit. BIOL 2001/PSYC 2002 Biological Dimensions of Human Sexuality: Emphasizes sex as a biological phenomenon. Considers important questions such as: What is sex? When did sex evolve? Why sex? Studies the history of sex and sex in nature. Serves as a source of information regarding what can be expected from the human body and modern issues concerning sexuality. Emphasizes human anatomy and physiology, human population, conception, pregnancy, birth, STDs, and AIDS. Spring, 3 hours lecture and discussion,

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 2001Biological Dimensions of Human Sexuality
Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 2003Lifecycle of Nutrition: Focuses on the3

fundamentals of normal nutrition from preconception to old age. Special attention is paid to the physiological principles and nutritional interventions that apply to each part of the life cycle. Applies case studies to real life at each stage in development, with consideration of cultural and socioeconomic influences on healthy eating behavior. Fall, 3 hours lecture and discussion,

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 2018Special Topics in Biology Tropical

Ecology & Conservation: Introduces students to key concepts in tropical ecology and conservation and provides hands-on experience collecting ecological data in a tropical country. Explores the ecology of tropical ecosystems, current issues in tropical conservation, and field methods to study

Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 2024Special Topics in Biology: Owl2

Prowl and Winter Raptors: Studies the wintering snowy owl and northern raptor locations in North America. The course includes a trip to Ontario, Canada. Not offered every year, Travel fee determined at time of trip. 2 hours lecture and discussion, field trip required,

Subject
BIOL
Credits (min)
2
Credits (max)
2
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 2110Pathophysiology

118 Keystone College

Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 2115Microbiology: Examines the4

importance and impact of microorganisms on the modern world with an emphasis on bacteria and viruses. Techniques for identifying microbes will be explored through hands-on experimentation with various microorganisms. and discussion, 3-hour laboratory,

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
“C” or better in BIOL 1125.
Corequisite
BIOL 2116. Fall, 3 hours lecture
BIOL 2120Introductory Ecology: Studies the4

natural assemblage of plants and animals. Emphasizes the interactions of both plant and animal populations in normally functioning ecosystems. Studies ecosystem structure, predation, competition, temperature, water, and energetic relationships; food webs, succession, and conservation biology. Laboratory focuses on field techniques, data analysis, and writing in acceptable scientific style.

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 1150 with a “C” or better. Corequisite: BIOL 2121. Spring and Intersession, 3 hours lecture and discussion, 3-hour laboratory,
BIOL 2135Science in Movies: Explore the3

science concepts presented in modern movies. Examines a group of modern movies and explores the feasibility of the science presented in each. Includes molecular biology, genetics, 201 Keystone College microbiology, virology, chemistry, physics, astronomy, environmental science, and geology. Explores ethics considerations. Fall and Spring, 3 hours lecture and discussion,

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 2140Introduction to Forensic Science II4

Builds upon the crime scene investigative sciences by introducing scientific crime laboratory techniques of various forensic disciplines including DNA analysis, toxicology, forensic chemistry, forensic anthropology, serology, questioned documents, and trace evidence.

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 1135 or consent of the instructor. and discussion, 3 hours laboratory,
Corequisite
BIOL 2141. Spring, 3 hours lecture
BIOL 2145Bioethics: Introduces theories and

concepts of ethics. Uses case studies, class discussion, and lecture to evaluate and analyze ethical problems and issues from various perspectives pertaining to the biological sciences. Students will be able to support ethical decisions with logical arguments. Prerequisites: ENGL 1110 and ENGL 1125, BIOL 1125 or CHEM 1120 or

Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 2150Dendrology: Introduces trees and3

other woody plant species that are native or commonly found in Pennsylvania and the region. Emphasizes native plants and covers other noteworthy species from other regions of the country and world. Emphasizes field identification and taxonomy.

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 1150. lecture and discussion,
Corequisite
BIOL 2151. Fall (even years), 3 hours
BIOL 2155Wildlife and Fisheries Biology
Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 2160Fall or Spring, lecture and3

discussion, 3 credits.

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
hours
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 2161Fall and Summer, lecture and3

discussion, 3 hours laboratory, 4 credits.

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
hours
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 2165Recommended elective for all3

students planning careers in human health. Fall or Spring (not offered every year), 3 hours lecture and discussion,

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 3002Nutrition: Studies the structural and3

functional relationships between biological molecules and human health. Studies the nutrients essential to human life and well-being. Studies nutrients relative to their function in metabolism, sources in food, and relationship to health. Fall, 3 hours lecture and discussion,

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 3010Special Topics in Biology Animal3

Behavior & Parasitism: An in-depth study and review of parasitic relationships and how they influence animal behavior. There are many connections between animal behavior and parasitism with several theories linking the two fields. Conceptual and technological advances have helped to determine new connections between animal behavior and the ecology and evolution of infectious diseases. Parasitism is a type of symbiotic relationship between a smaller parasite and the larger host. In order to survive, parasites and hosts compete with each other in an evolutionary tug-of-war to try to outwit the other. Parasites seek to enhance infectivity, and hosts strive to remain unparasitized. Examines the relationships between hosts and parasites including the physical and physiological mechanisms utilized by each group.

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 1125 and 1130. Fall or Spring (not offered every year), 3 hours lecture, discussion, hands-on assignments,
BIOL 3110Cell Biology: Studies the structure4

and function of a cell and its subcellular organelles. Focuses on biological macromolecules, enzyme, biological transport, gene expression, motility, and new technologies.

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 1125. hours lecture and discussion, 3 hours laboratory,
Corequisite
BIOL 3111. Fall (odd years), 3
BIOL 3115Wetland and Riparian Ecology

Focuses on major concepts in wetland ecology including wetland definitions, identification and delineation, and wetlands functions and values. Discusses wetland classification systems; state and federal jurisdiction and policies; and regulatory issues in wetland conservation, protection, and management. Includes field trips to on- and off-campus sites. at time of travel. Fall (odd years), 3 hours lecture and discussion, 3 credits. 202 Keystone College

Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 2120. Travel fee determined
BIOL 3135Limnology: Investigates lakes4

reservoirs, ponds, and streams as dynamic systems. Considers the physical, chemical, geological, and biological components of aquatic ecosystems. Emphasizes quantitative sampling and analytical techniques.

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 2120, CHEM 1120 and CHEM/ENVT 3125. lecture and discussion, 3 hours laboratory,
Corequisite
BIOL 3136. Fall (even years), 3 hours
BIOL 3150Virology: Covers a detailed survey of3

viruses that are important to both humans and animals. Topics include classification, structure, penetration and replication techniques, immune system responses, vaccines, diagnostic techniques, bioterrorism, and pandemic spread. Spring (odd years), 3 hours lecture, discussion, and demonstrations,

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 1125 and BIOL 2115.
BIOL 3160Immunology: Designed for upper3

level biology majors. Covers the study of the basic principles of immunology, focusing on the immune system and how this system protects the body from infectious agents.

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 1125, BIOL 2160, and BIOL 2165. Spring (even years), 3 hours lecture and discussion,
BIOL 3170Genetics: Emphasizes genetic and3

evolutionary mechanisms, gene interaction, polygenic inheritance, selection, mutation, human diversity, human mating systems, management of the human gene pool, and the direction of present and future research. hours lecture and discussion,

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 1125 and BIOL 1130. Spring, 3
BIOL 3175Ichthyology: Presents an intense

study of fish. Exposes students to the range of natural history, ecological, and management concepts, as well as receiving hands-on field training. Features active lectures and introduces a research project to the taxonomic structures in intensive laboratories. Spends class time on discussing the evolution, behavior, physiology, and conservation of fish. Through assorted readings and field experiences, covers the evolution and ecology of the different taxa within these groups. Allows students to gain an appreciation for and identify the fish diversity in Pennsylvania and throughout North America. Field trips serve as a means to underscore and/ or illustrate the concepts discussed in class, as well as an understanding of techniques used to safely work

Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 3180Ethics and Quality in Forensic3

Science: Builds upon students’ scientific background in forensics by focusing on understanding ethical issues encountered by forensic scientists and quality programs. Use case studies, class discussion, and lecture to evaluate and analyze ethical problems and illustrate applications to quality systems in a forensic laboratory. Examines codes of ethics, professionalism, obligations of the forensic witness, bias, quality assurance and quality control, and report writing. Participate in role playing, discussions, debates, writing exercises, and presentations throughout the course.

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 1135 or BIOL 2140. Spring (not offered every year), 3 hours lecture and discussion,
BIOL 3185Cancer Biology: Provides the3

fundamentals for understanding what cancer is and how to treat it. Lectures focus on how genetic modifications with molecular altering consequences lead to development and spread of cancer cells and how we diagnose and treat the disease. Covers oncogenes, tumor suppression, tumorigenesis, angiogenesis, metastasis, and classical as well as future treatments of cancer.

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 3110 or BIOL 3170. Fall or Spring (not offered every year), 3 hours lecture and discussion,
BIOL 3190Comparative Vertebrate Anatomy4

Emphasizes comparative development, functional anatomy and macroevolution of vertebrate body plans. Examines the diversity and phylogenetic history of fossil and local forms, development and comparative embryology of each organ system, and comparative functional anatomy of the major clades of living vertebrates within an evolutionary framework.

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 1130. Corequisite: BIOL 3191. Fall, 3 hours lecture and discussion, 3 hours laboratory,
BIOL 3195Evolution: Explores the theory of3

evolution from the early generation of the theory to the evidence present today. Discusses the mechanisms for evolutionary change and 203 Keystone College adaptation within and between species. Analyzes evolutionary evidence.

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 1130. Spring (even years), 3 hours lecture and discussion,
BIOL 3200General Physiology: Examines body4

system functions from the cellular level through the organismal level of organization. Special emphasis placed on the important linkage between the cardiovascular, respiratory, endocrine, and excretory systems.

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 2165 or BIOL 3190, and CHEM 1125. Corequisite: BIOL 3201. Spring (even years), 3 hours lecture and discussion, 3 hours laboratory,
BIOL 3210Principles of Biotechnology: Looks4

at the principles, techniques, and applications of biotechnology. Discusses specific techniques such as recombinant DNA technologies, protein purification, and large-scale industrial harvesting of products. Examines bioremediation and drug discovery. Discusses the processes that take an idea from development to a marketable product. 3211. Spring (not offered every year), 3 hours lecture and discussion, 3 hours laboratory,

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 3240. Corequisite: BIOL
BIOL 3215Microbial Genetics: Explores in detail4

the genetic workings of the microbial world. Examines topics such as gene expression, DNA repair, and gene transfer mechanisms including plasmids, transposons, and transduction. Discusses how we, as humans, are entering the “-omic” age of discovery in biology and related topics such as genomics, proteomics, and metabolomics.

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 2115. hours lecture and discussion, 3 hours laboratory,
Corequisite
BIOL 3216. Spring (odd years), 3
BIOL 3220Microbial Physiology: Explores in4

detail processes, including biochemical, molecular, and cellular, that bacteria use to survive and cause disease. The understanding of these pathways is fundamental in several areas of study including microbiology, immunology, and virology. These pathways are utilized and exploited in the food, pharmaceutical, medical and biotechnological industries. Knowledge of these processes aid in the understanding and preventing of disease and food spoilage.

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 2115. hours lecture and discussion, 3 hours laboratory,
Corequisite
BIOL 3221. Spring (even years), 3
BIOL 3225Mammalogy: An intense study of

mammals. Exposes students to the range of natural history, ecological, and management concepts, as well as receiving hands-on field training. Features active lectures and a research project, and introduces the taxonomic structures in intensive laboratories. Discusses the evolution, behavior, physiology, and conservation of mammals. Through assorted readings and field experiences, explores the evolution and ecology of the different taxa within these groups. Develops students’ appreciation for and identification of the mammal diversity in Pennsylvania and throughout North America. Field trips serve as a means to underscore and/ or illustrate the concepts discussed in class, as well as an understanding of techniques used to safely work with mammals in the field. hours lecture and discussion, 3 hours laboratory, field trips (travel fee determined at time of travel), 4 credits.

Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 2120. Fall (odd years), 3
BIOL 3230Ornithology: An intense study of

birds. Exposes students to the range of natural history, ecological, and management concepts, as well as hands-on field training. Features active lectures and a research project and introduces the taxonomic structures in intensive laboratories. Discusses the evolution, behavior, physiology, and conservation of birds. Examines evolution and ecology of the different taxa within these groups through assorted readings and field experiences. Students gain an appreciation for and learn to identify the bird diversity in Pennsylvania and throughout North America. Field trips serve as a means to underscore and/or illustrate the concepts discussed in class, as well as an understanding of techniques used to safely work with birds in the field. Prerequisite: BIOL 2120. Spring (odd years), 3 hours lecture and discussion, 3 hours laboratory, field trips (travel fee determined at time of travel), 4 credits.

Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 3235Herpetology: An intense study of4

reptiles and amphibians. Exposes students to the range of natural history, ecological, and management concepts, as well as receiving hands-on field training. Features active lectures and a research project and introduces the taxonomic structures in intensive laboratories. Discusses the evolution, behavior, physiology, and conservation of reptiles and amphibians. Examines, through assorted readings and field experiences, evolution and ecology of the 204 Keystone College different taxa within these groups. Advances students’ appreciation for and identification of reptile and amphibian diversity in Pennsylvania and throughout North America. Field trips serve as a means to underscore and/or illustrate the concepts discussed in class, as well as an understanding of techniques used to safely work with reptiles and amphibians in the field. lecture and discussion, 3 hours laboratory, field trips (travel fee determined at time of travel), 4 credits. BIOL/CHEM 3240 Biochemistry I: Studies the major classes of biological molecules including carbohydrates, lipids, amino acids, proteins, and nucleic acids. Examines how structure and function is interrelated and how poor structure will lead to a disease state. Prerequisites: CHEM 2110 and CHEM 2115 with a grade of “C” or better. Fall, 3 hours lecture and discussion, 3 hours laboratory. BIOL/CHEM 3245 Biochemistry II: Examines the metabolic pathways of life, including the anabolic and catabolic pathways for carbohydrates, lipids, proteins, amino acids, and nucleic acids. Reviews enzyme structure and function since these are important and necessary components of any metabolic pathway. Prerequisite: “C” or better in BIOL/CHEM 3240. Spring, 3 hours lecture and discussion, 3 credits.

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 2120. Fall (odd years), 3 hours
BIOL 3240Biochemistry I

BIOL/CHEM 3245 – Biochemistry II BIOL 4810 /ENVT4810– Seminar and Research

Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 3250Entomology: This course is presented4

as an intense study of insects. Students will be exposed to the range of natural history, ecological, and management concepts, as well as receiving hands-on field training. Students will have active lectures and a research project and will be introduced to the taxonomic structures in intensive laboratories. Class time will be spent discussing the evolution, behavior, physiology, and conservation of insects. Through assorted readings and field experiences, students will learn about the evolution and ecology of the different taxa within these groups. Further, students will be able to gain an appreciation for and identify the insect diversity in PA and throughout North America. Field trips will serve to underscore and/or illustrate the concepts discussed in class, as well as an understanding of techniques used to safely work with insects in the field.

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 1125 and 1130. Corequisite: BIOL 3251. Fall (not offered every year), 3 hours lecture and discussion, 3 hours laboratory,
BIOL 4110Forensic Microscopy and Trace4

Analysis: Focuses on furthering students’ ability to use a microscope and analytical instrumentation to analyze various types of evidence that may be encountered in a forensic investigation. Examines optics and how microscopes work that are commonly used in forensic trace evidence analysis, as well as common types of trace evidence (hairs, fibers, glass, etc.). Emphasizes students critically analyzing literature involving new methods of trace evidence analysis and presenting it to their peers. At the completion of the course, it is expected that students’ will have a thorough knowledge of microscopes, how trace evidence is analyzed, and have advanced their laboratory skills to the analysis of unknown trace evidence. 4111. Spring (alternating years), 3 hours lecture and discussion. 3 hours laboratory,

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
CHEM 1125. Corequisite: BIOL
BIOL 4115Molecular Biology: Furthers the3

students’ understanding of the fundamentals of modern molecular biology. Covers DNA and RNA, gene expression, epigenetics, recombinant DNA, and biotechnology. Emphasizes students critically analyzing the molecular biology literature and applying this knowledge to experimental design. At the completion of the course, it is expected that students will have a thorough knowledge of molecular biology mechanisms, interpreting experimental data, and how to design experiments to answer questions in molecular biology.

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 3170. Fall (even years), 3 hours lecture and discussion,
BIOL 4120Conservation Biology: Explores3

the role of science in the conservation of animals and plants through lectures, text readings, case studies, and readings from the primary literature. Lectures focus on an introduction to the scientific study and analysis of biodiversity; a survey of the major threats to biodiversity; an overview of contemporary conservation strategies and an analysis of recent and historic success and failures in conservation efforts.

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 2120. Spring (odd years), 3 hours lecture and discussion,
BIOL 4125Forensic Biology: Explores4

fundamental procedures of forensic DNA testing and bodily fluid identification. Focuses on furthering the students’ knowledge of genetics and physiology of the human body by applying it 205 Keystone College to forensic investigations. Covers the identification of common bodily fluids (semen, blood, saliva), novel methods and research in body fluid identification and DNA analysis (next generation sequencing), and current DNA analysis technologies. Emphasizes students critically analyzing literature involving new methods of forensic biology testing. At the completion of the course, students should have a thorough knowledge of forensic biology, how biological evidence is processed, and have advanced their laboratory skills for biological evidence testing. discussion, 3 hours laboratory,

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 3170.
Corequisite
BIOL 4126. Fall, 3 hours lecture and
BIOL 4130Microbial Ecology: Reviews how4

microorganisms interact with each other and other living and nonliving components of their environments. Examines the role microorganisms play in important environmental cycles such as carbon, nitrogen, and other biomolecules. Students examine how to evaluate microbial diversity and identify applications of microbial ecology. Covers what effect climate change might have on this microbial diversity. years), 3 hours lecture and discussion, 3 hours laboratory,

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 2115 and BIOL 2120. Fall (odd
BIOL 4135Microbial Physiology
Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 4140Forensic Validation: Explores a2

fundamental process in forensic DNA testing and bodily fluid identification – method validation. Focuses on furthering the students’ knowledge of genetics and physiology of the human body by studying a method in depth. Topics to discuss include bodies that recommend validation and its procedures, experimental design, report writing, statistics, and critical analysis of data. Emphasizes students critically analyzing their own results from using new methods. At the completion of the course, it is expected that students will have a thorough knowledge of forensic validation, how a validation is conducted, and have advanced their laboratory skills for biological evidence testing. The final product of the course is a written validation report.

Subject
BIOL
Credits (min)
2
Credits (max)
2
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 3170 and MATH 2115. Spring (alternating years), 2 hours lecture and laboratory,
BIOL 4145Animal Behavior

Botany (6 credits minimum) Field Biology/Science Skills (maximum 2 credits chosen from 1023, 1030, 1052, 1077; SCI 1010, 1013)

Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 4150Mycology: Examines a broad4

survey of the Kingdom Fungi. Explores molecular systematics, physiology, cellular organization, reproduction and dispersal, roles of fungi in ecosystems, fungal symbioses, plant and animal pathogens, predaceousfungi, biological control of fungi, fungal food fermentation, mycotoxins and food spoilage, pharmaceuticals, and fungus-insect/fungus- insect-plant interactions.

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 1150. Corequisite: BIOL 4151. Fall (even years), 3 hours lecture and discussion, 3 hours laboratory,
BIOL 4155Wildlife Management: To kill or not3

to kill, that is the question. Hunting in North America today is a decidedly different enterprise than that which our forebears practiced even up to one hundred years ago. Hunting has shifted from a practice borne out of utilitarian necessity to an endeavor based on choice and leisure preference. It has grown from a rural chore to a full-fledged recreational industry with superstores, luxury guided vacation packages, and a full slate of television and online programming. Examines managing game species, their habitats, and those that pursue them, and the history behind hunting and hunting culture.

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 2120. Spring (even years), 3 hours lecture and discussion,
BIOL 4165Ecological Restoration

139 Keystone College 33 BIOL 1015, 1038, BIOL 1013, 1014,

Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 4710Biology Internship Preparation

Prepares students for the identification, placement, and successful completion of an internship in the biological science field. Fall, Spring or Summer, 0 credits. Open only to degree seeking students.

Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 4715Biology Internship I: Provides3

supervised exposure to the work environment and practical experience through an internship at one or more pre-approved sites. Fall, Spring or Summer, 40 hour per credit. 1-6 credits. Open only to degree seeking students. BIOL 4810/ENVT 4810 Biology Seminar and Research: Begins the capstone research process. Students propose their senior capstone research project, prepare an extensive literature review of the topic, and prepare for graduate school or job interviews. Students MUST have an approved research proposal to continue to BIOL 4910: Capstone Research Project.

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
Senior status. Fall, 3 hours lecture and discussion,
BIOL 4810Biological Seminar and Research
Subject
BIOL
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
BIOL 4910Capstone Research Projects4

Provides a research project under the direction of a faculty member. Research hours will be determined based on agreement between student and faculty mentor. A presentation of the research project will be made to the campus community in the form of a written and/or oral presentation of findings.

Subject
BIOL
Credits (min)
4
Credits (max)
4
Credit unit
credits
Type
course
Edition
2025-2026-undergraduate
Source
www.keystone.edu
Prerequisite
BIOL 4810 with a “C” or better AND senior status AND approval of instructor and mentor. Spring, 3 hours lecture and discussion, 3 hours laboratory,
BIOL 5100Scientific Writing and Communication

Animal & Natural Resource Management Elective Animal & Natural Resource Management Elective Ecology and Organismal Biology Elective Ecology and Organismal Biology Elective Ecology and Organismal Biology Elective Ecology and Organismal Biology Elective Environmental and Physical Science Elective Laboratory Courses (choose three courses from the list below):

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5150Stream Ecology: Covers the principles of stream, river, and watershed ecology. Applies

these principles to the understanding of the natural history and ecology of running waters. Examines the composition and function of biota in streams and rivers as well as the interactions of that biota with their physical and chemical environments. The course will address human influences on these ecosystems. 3 credits. Every other Spring.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5151Stream Ecology Lab: Students apply and study the principles of stream, river, and watershed

ecology. Examines the physical, chemical and biological qualities of waterways. This course is expected to be taken either concurrently or following BIOL 5150 Stream Ecology. 1 credit. Every other Spring.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5155Ecological Restoration: This course introduces students to the fundamental principles of

restoration ecology, the science of restoring degraded ecosystems. Students will explore the causes of ecosystem degradation, methods and motivations for restoration, and how to evaluate restoration success. This course will also consider the social, political, and cultural dimensions of ecological restoration. 3 hours lecture and discussion, 3 hours laboratory. 3 credits. Every other Spring.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5156Ecological Restoration Lab: This laboratory provides hands-on experience working with

ecological restoration techniques and also develops skills in soil and water testing, invasive species identification, and horticultural practices. Students will also consider how the human dimensions of restoration influence their work. 1 credit. Every other Spring.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5157Tropical Ecology Lab: This laboratory introduces students to key concepts in tropical

ecology and provides hands-on experience with field methods to study plants and animals in a tropical ecosystem. Features a week-long field trip where students will experience field work in a tropical country and will work with local researchers and conservation groups. The focal organisms and country will vary with each course offering. Travel fee is determined at the time of travel. 1 credit. Offered at the discretion of the department. 33 Keystone College Graduate Catalog

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5160Forest Ecology: Allows students to gain an understanding of forest ecosystem structure

function, composition, and dynamics and will apply ecological principles to current environmental problems impacting forests. 3 credits. Every other Fall.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5161Forest Ecology Lab: Provides firsthand experience conducting qualitative and quantitative

measurements within a forest ecosystem and analyzing that data. Students will apply their experiences to understanding how forest stands change over time and how that affects biota within the ecosystem. 1 credit. Every other Fall.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5165Environmental Toxicology: Introduces students to the principles of environmental toxicology

and the major classes of pollutants. Discusses the biological effects of environmental toxicants on terrestrial as well as aquatic wildlife. 3 credits. Offered at the discretion of the department.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5170Aquatic Macroinvertebrate Zoology: Students will gain an understanding of the

classification, structure, and life history of lotic and lentic freshwater macroinvertebrates. A portion of the course emphasizes Pennsylvania species, specifically those that might be found in the Howard Jennings Nature Preserve or local area waters. 3 credits. Offered at the discretion of the department.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5171Aquatic Macroinvertebrate Zoology Lab: Students gain hands on experience in the

collection and identification of freshwater macroinvertebrates. This lab portion of the course will place an emphasis on Pennsylvania species, specifically those that might be found in the Howard Jennings Nature Preserve or local area waters. 1 credit. Offered at the discretion of the department.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5175Ecology of Wildlife Populations: Introduces students to concepts of animal ecology as it

applies to various levels of organization: ecosystem, community, population and individual. Students investigate the mechanisms that impact wildlife populations and how ecological principles can be applied in wildlife conservation. 3 credits. Offered at the discretion of the department.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5180Fish Management and Culture: Introduces students to the principles of fisheries science and

management. Provides an overview of the processes and tools for managing fish populations and their habitats as well as emphasizing quantitative methods of fisheries assessment. 3 credits. Every other Spring.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5185Natural Resources Policy and Management

31 Keystone College Graduate Catalog

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5190Population Dynamics: Presents the theory necessary for understanding wildlife population

dynamics and explains how to use theory and data to inform management and conservation efforts. 3 credits. Offered at the discretion of the department.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5195Aquatic Ecosystem Conservation: Provides students with an understanding of human

impacts in freshwater ecosystems and the challenges and approaches for conserving freshwater diversity and ecosystem services. 3 credits. Offered at the discretion of the department.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5200Microbial Ecology: Reviews how microorganisms interact with each other and other living

and nonliving components of their environments. Examines the role microorganisms play in important environmental cycles such as carbon, nitrogen, and other biomolecules. Students examine how to evaluate microbial diversity and identify applications of microbial ecology. 3 credits. Offered at the discretion of the department. 34 Keystone College Graduate Catalog

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5210Animal Behavior: Examines the development of major areas in conservation-oriented

research that include patterns of management, biodiversity, extinction, conservation genetics, conservation of populations, communities and landscapes, and ecological sustainability. 3 credits. Every other Spring.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5215Mammalogy: Presented as an intense study of mammals. Explores students to the range of

natural history, ecological, and management concepts, as well as hands-on field training. Provides students with active lectures and a research project and introduces the taxonomic structures in intensive laboratories. Discusses the evolution, behavior, physiology, and conservation of mammals. Examines the evolution and ecology of the different taxa within these groups through assorted readings and field experiences. Allows students to gain an appreciation for and identify the mammal diversity in PA and throughout North America. Field trips will serve to underscore and/or illustrate the concepts discussed in class, as well as an understanding of techniques used to safely work with mammals in the field. 3 credits. Every other Fall.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5216Mammalogy Laboratory: This laboratory course is presented as an intense study of

mammals. Students will be exposed to the range of natural history, ecological, and management concepts, as well as receiving hands-on field training. Students will have active lectures and will be introduced to the taxonomic structures in intensive laboratories. Class time will be spent discussing the evolution, behavior, physiology, and conservation of mammals. Through assorted readings and field experiences, students will learn about the evolution and ecology of the different taxa within these groups. Further, students will be able to gain an appreciation for and identify the mammal diversity in PA and throughout North America. Field trips will serve to underscore and/or illustrate the concepts discussed in class, as well as an understanding of techniques used to safely work with mammals in the field. 3 hours laboratory and field trips. 1 credit. Every other Fall.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5220Ornithology: Presented as an intense study of birds. Exposes students to the range of

natural history, ecological, and management concepts, as well as receiving hands-on field training. Features active lectures and a research project and introduces the taxonomic structures in intensive laboratories. The evolution, behavior, physiology, and conservation of birds. Students learn about the evolution and ecology of the different taxa within these groups through assorted readings and field experiences. Provides students with an appreciation for and identifying bird diversity in PA and throughout North America. Field trips serve to underscore and/or illustrate the concepts discussed in class, as well as an understanding of techniques used to safely work with birds in the field. 3 credits. Every other Spring.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5221Ornithology Laboratory: This course is presented as an intense study of birds. Students will

be exposed to the range of natural history, ecological, and management concepts, as well as receiving hands-on field training. Students will have active lectures and a research project and will be introduced to the taxonomic structures in intensive laboratories. Class time will be spent discussing the evolution, behavior, physiology, and conservation of birds. Through assorted readings and field experiences, students will learn about the evolution and ecology of the different taxa within these groups. Further, students will be able to gain an appreciation for and identify the bird diversity in PA and throughout North America. Field trips will serve to underscore and/or illustrate the concepts discussed in class, as well as an understanding of techniques used to safely work with birds in the field. 3 hours laboratory. 1 credit. Every other Spring.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5225Ichthyology: An intense study of fish that exposes students to the range of natural history

ecological, and management concepts through hands-on field training. Features active lectures and a research project and introduces the taxonomic structures in intensive laboratories. Discusses the evolution, behavior, physiology, and conservation of fish. Examines the evolution and ecology of the different taxa within these groups through assorted readings and field experiences. Allows students to 35 Keystone College Graduate Catalog

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5226Ichthyology Laboratory: This course is presented as an intense study of fish. Students will

be exposed to the range of natural history, ecological, and management concepts, as well as receiving hands-on field training. Students will have active lectures and a research project and will be introduced to the taxonomic structures in intensive laboratories. Class time will be spent discussing the evolution, behavior, physiology, and conservation of fish. Through assorted readings and field experiences, students will learn about the evolution and ecology of the different taxa within these groups. Further, students will be able to gain an appreciation for and identify the fish diversity in PA and throughout North America. Field trips will serve to underscore and/or illustrate the concepts discussed in class, as well as an understanding of techniques used to safely work with fish in the field. 3 hours laboratory. 1 credit. Every other Fall.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5230Herpetology: An intense study of reptiles and amphibians that exposes students to the range

of natural history, ecological, and management concepts through hands-on field training. Features active lectures and a research project and introduces the taxonomic structures in intensive laboratories. Discusses the evolution, behavior, physiology, and conservation of reptiles and amphibians. Examines evolution and ecology of the different taxa within these groups through assorted readings and field experiences. Allows students to gain an appreciation for and identify the bird diversity in PA and throughout North America. Field trips serve to underscore and/or illustrate the concepts discussed in class, as well as an understanding of techniques used to safely work with reptiles and amphibians in the field. 3 credits. Every other Fall.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5231Herpetology Laboratory: This laboratory course is presented as an intense study of reptiles

and amphibians. Students will be exposed to the range of natural history, ecological, and management concepts, as well as receiving hands-on field training. Students will have active lectures and a research project and will be introduced to the taxonomic structures in intensive laboratories. Class time will be spent discussing the evolution, behavior, physiology, and conservation of reptiles and amphibians. Through assorted readings and field experiences, students will learn about the evolution and ecology of the different taxa within these groups. Further, students will be able to gain an appreciation for and identify the reptile and amphibian diversity in PA and throughout North America. Field trips will serve to underscore and/or illustrate the concepts discussed in class, as well as an understanding of techniques used to safely work with reptiles and amphibians in the field. 3 hours laboratory. 1 credit. Every other Fall.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5235Wetland and Riparian Ecology: Focuses on major concepts in wetland ecology including

wetland definitions, identification of habitat types, and wetlands functions and values. Discusses wetland classification systems; state and federal jurisdiction and policies; and regulatory issues in wetland conservation, protection, and management. Studies wetland delineation and various assessment techniques. Includes field trips to on and off campus sites. 3 credits. Every other Fall.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5240Wildlife Management: To kill or not to kill, that is the question. Hunting in North America

today is a decidedly different enterprise than that which our forebears practiced even up to one hundred years ago. Hunting has shifted from a practice borne out of utilitarian necessity to an endeavor based on choice and leisure preference. It has grown from a rural chore to a full-fledged recreational industry with superstores, luxury guided vacation packages, and a full slate of television and online programming. Examines managing game species, their habitats, and those that pursue them, and the history behind hunting and hunting culture. 3 credits. Every other Spring. 36 Keystone College Graduate Catalog

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5245Conservation Biology

Environmental and Physical Sciences:

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5250Limnology: Investigates lakes, reservoirs, ponds, and streams as dynamic systems

Considers the physical, chemical, geological, and biological components of aquatic ecosystems. Emphasizes quantitative sampling and analytical techniques. 3 credits. Every other Fall.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5251Limnology Laboratory: This course investigates lakes, reservoirs, ponds, and streams as

dynamic systems. Considers the physical, chemical, geological, and biological components of aquatic ecosystems and emphasizes quantitative sampling and analytical techniques as they are applied to aquatic ecosystems. 3 credits. Every other Fall.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5255Soils Science

Analytical Tools, Research, and Statistical Analysis Elective:

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5260Geospatial Methods in Resource Management

32 Keystone College Graduate Catalog

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5265Mycology: Examines a broad survey of the Kingdom Fungi. Explores molecular systematics

physiology, cellular organization, reproduction and dispersal, roles of fungi in ecosystems, fungal symbioses, plant and animal pathogens, predaceous fungi, biological control of fungi, fungal food fermentation, mycotoxins and food spoilage, pharmaceuticals, and fungus-insect/fungus-insect- plant interactions. 3 credits not offered every year. Every other Fall.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5266Mycology Laboratory: Students gain hands on experience in the identification of fungi and

their structures, as well as preservation and laboratory culture techniques. Additionally, the role of fungi in ecosystems and in human society will be explored. This lab portion of the course will place an emphasis on Pennsylvania species, specifically those that might be found on the Keystone College campus and surrounding areas. 3 hours laboratory. 1 credit. Every other Fall.

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5270Animal Behavior and Parasitism: An in-depth study and review of parasitic relationships

and how they influence animal behavior. There are many connections between animal behavior and parasitism with several theories linking the two fields. Conceptual and technological advances have helped to determine new connections between animal behavior and the ecology and evolution of infectious diseases. Parasitism is a type of symbiotic relationship between a smaller parasite and the larger host. To survive, parasites and hosts compete in an evolutionary tug-of-war to try to outwit the other. Parasites seek to enhance infectivity, and hosts strive to remain unparasitized. Examines the relationships between hosts and parasites including the physical and physiological mechanisms utilized by each group. 3 hours lecture and discussion. 3 credits. Offered at the discretion of the department. 37 Keystone College Graduate Catalog

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5275Entomology

Animal and Natural Resource Management Electives:

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 5276Entomology Laboratory

And choose Thesis or Non-Thesis Track

Subject
BIOL
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 6810Biological Thesis Research I: This course is intended for Master level students to begin or3

continue their thesis research. This course provides students with the time to actively work on their research, data analysis, and writing. Students will work with their graduate program advisor on successful progression of their research throughout the semester. Successful completion of this course is indicated by observable progress on the student’s thesis research.

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu
BIOL 6910Biological Thesis Research II: This course is intended for Master level students to pursue3

the completion of their thesis research. This course provides students with the time to actively work on their research, data analysis, and writing. Students will work with their graduate program advisor on successful progression of their research throughout the semester. Successful completion of this course is indicated by the completion of their research, the public presentation or their research, and readiness for their thesis defense.

Subject
BIOL
Credits (min)
3
Credits (max)
3
Credit unit
credits
Type
course
Edition
2025-2026-graduate
Source
www.keystone.edu

Source: Keystone College's catalog, linked per course · table learning_unit · CourseShelf publish 59